Euphorbia exstipulata |
Euphorbia acuta |
|
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square-seed spurge, squareseed or Clark Mountain spurge |
point sandmat |
|
Habit | Herbs, annual, with slender taproot. | Herbs, perennial, with strongly thickened rootstock. |
Stems | erect, 5–26 cm, uniformly puberulent to hispidulous or glabrous; branches arcuate. |
ascending to erect, 5–30 cm, uniformly and densely canescent or sericeous. |
Leaves | opposite; petiole 1–3 mm, often indistinct, glabrous or puberulent; blade linear to narrowly elliptic or ovate, 14–42 × 3–28 mm, base attenuate, margins coarsely serrate, occasionally revolute, apex acute to obtuse, abaxial surface sparsely hispidulous to strigillose, adaxial surface glabrous; midvein conspicuous. |
opposite; stipules deciduous, sometimes appearing absent, distinct, brown, linear-subulate, thin, 0.3–0.8 mm, canescent; petiole 0.4–1.2 mm, moderately to densely canescent; blade ovate to lanceolate, 6–20 × 3–8 mm, base subsymmetric, rounded to cuneate, margins entire, strongly involute, apex long-acuminate, spinulose, abaxial surface canescent to densely sericeous, adaxial surface glabrous or sparsely canescent; 3-veined from base but only midvein conspicuous. |
Involucre | turbinate to campanulate, 1.1–1.5 × 1–1.3 mm, glabrous, pilose or puberulent; involucral lobes divided into several linear lobes; glands 4(–5), yellow to pink, sessile and broadly attached, 0.2 × 0.3–0.4 mm, opening oblong to nearly circular, glabrous; appendages usually petaloid, white to pink, ovate to trapezoidal, occasionally absent, not incurved and covering glands, 0.2–0.4 × 0.3–0.8 mm, entire, undulate, or conspicuously divided into triangular segments, glabrous. |
turbinate to urceolate, 2–2.6 × 1.7–2.5 mm, villous to lanate; glands 4, yellow-green to orange or red, slightly concave, oblong-elliptic, 0.2–0.4 × 0.6–1.5 mm; appendages white, flabellate, 1.1–2.1 × 0.2–0.6 mm, distal margin shallowly and irregularly toothed. |
Staminate flowers | 10–12. |
20–25. |
Pistillate flowers | ovary puberulent on keels, styles 0.8–1.1 mm, 2-fid 1/2 to nearly entire length. |
ovary strigose, pubescent to villous; styles 0.6–0.9 mm, 2-fid 1/2 length. |
Capsules | broadly depressed-oblong to ovoid, 2.7–3.3 × 3.1–3.9 mm, puberulent (with appressed hairs usually concentrated on keels); columella 1.9–2.5 mm. |
subglobose to broadly ovoid, 2.8–3.7 mm diam., strigose, pubescent to villous; columella 2.3–3 mm. |
Seeds | white to gray or light brown, ovoid, bluntly 4-angled in cross section, 1.9–2.5 × 1.4–1.7 mm, tuberculate, often with 2 transverse ridges; caruncle 0.1 × 0.2 mm. |
white, ovoid, 4-angled in cross section, 2.2–2.6 × 1.1–1.4 mm, smooth to finely reticulate. |
Cyathial | arrangement: terminal cymose or dichasial branches usually 1–2, occasionally reduced to monochasia, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts similar in shape to distal leaves but smaller or highly reduced. |
|
Cyathia | peduncle 1–1.9 mm. |
solitary at distal nodes; peduncle 1.3–3.2 mm. |
2n | = 28, 48, 56. |
|
Euphorbia exstipulata |
Euphorbia acuta |
|
Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting spring–fall. |
Habitat | Desert scrub, grasslands, mesquite savannas, oak and oak-juniper woodlands. | Desert scrub, grasslands, oak-juniper savannas, limestone, rocky, sandy, or clay soils. |
Elevation | 800–2000 m. (2600–6600 ft.) | 400–1900 m. (1300–6200 ft.) |
Distribution |
AZ; CA; NM; TX; UT; WY; Mexico (Chihuahua, Coahuila, Durango, Guanajuato, San Luis Potosí, Sonora, Zacatecas)
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NM; TX; Mexico (Chihuahua, Coahuila) |
Discussion | Euphorbia exstipulata is native from Texas to California and northern Mexico. The species was found once in the late nineteenth century in Wyoming but has not been re-collected there. Broad-leaved plants have been segregated as var. lata, but the variation in leaf shape is continuous and no varieties are formally recognized here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia acuta is easily distinguished in the field by its relatively large, strongly involute, hairy, and acutely pointed leaves. The name Euphorbia acuta Engelmann has been proposed for conservation against the earlier name E. acuta Bellardi ex Colla (P. E. Berry et al. 2011). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 322. | FNA vol. 12, p. 259. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. exstipulata var. lata | Chamaesyce acuta, E. acuta var. stenophylla, E. georgei |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) — name proposed for conservation |
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