Euphorbia exserta |
Euphorbia georgiana |
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coastal sand spurge, maroon or purple sand spurge |
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Habit | Herbs, perennial, with spreading rootstock. | Herbs, annual, with taproot. |
Stems | erect or ascending, unbranched or branched, solitary, few, or occasionally densely clumped, previous year's dead stems not persistent, 20–33 cm, usually glabrous, rarely sparsely villous, glaucescent. |
erect, often branched near base, 10–18 cm, glabrous. |
Leaves | alternate; stipules less than 0.1 mm; petiole (0–)1–3 mm, glabrous; blade linear or linear-elliptic to obovate or orbiculate, proximal greatly reduced, scalelike, 15–30 × 1–20 mm, base cuneate to rounded, margins entire, apex rounded, broadly acute, or emarginate, surfaces glabrous, glaucescent; venation often obscure on narrow leaves, midvein conspicuous. |
petiole absent; blade oblanceolate, 5–12 × 3–5 mm, base attenuate, margins entire, apex rounded, surfaces glabrous; venation pinnate, midvein prominent. |
Involucre | usually dark red, campanulate, 1.3–1.6 × 1.4–2.1 mm, glabrous; glands 5, usually dark red, rarely greenish red, elliptic reniform, thickened, 0.3–0.5 × 0.8 mm; appendages white or green, often forming narrow rim around distal margin of gland, 0–0.2 mm, entire. |
infundibular, 1–1.1 × 0.4–0.5 mm, glabrous; glands 4, crescent-shaped; 0.3–0.4 × 0.4–0.5 mm; horns divergent, 0.3–0.5 mm. |
Staminate flowers | 20–25. |
5–10. |
Pistillate flowers | ovary glabrous; styles 0.4–0.8 mm, 2-fid at apex. |
ovary glabrous; styles 0.5 mm, 2-fid. |
Capsules | depressed-globose, 1.8–2.5 × 3.6–4.4 mm, glabrous; columella 1.9–2.4 mm. |
depressed-ovoid, 2.2–2.4 × 3.2–3.4 mm, 3-lobed; cocci rounded, smooth, glabrous; columella 2–2.1 mm. |
Seeds | ashy white, ovoid, 2.1 × 1.3 mm, angled with 5 blunt longitudinal ridges, with shallow and irregular pits; caruncle absent. |
gray, ovoid, 1.6–1.7 × 1.4–1.6 mm, with deep, rounded pits irregularly scattered over entire surface; caruncle reniform, subconic, 0.5–0.6 × 0.6–0.8 mm. |
Cyathia | usually in terminal dichasia, sometimes pleiochasia; peduncle 6–33 mm, filiform, glabrous. |
peduncle 0.3–0.5 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3, 1–3 times 2-branched; pleiochasial bracts rotund-obovate, wider than distal leaves; dichasial bracts distinct, not imbricate, broadly deltate to subreniform, base truncate to emarginate, margins entire, apex rounded to bluntly acuminate; axillary cymose branches 0–1. |
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Euphorbia exserta |
Euphorbia georgiana |
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Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting spring. |
Habitat | Xeric to dry pine-oak scrub of sand hills, pine-oak woodlands, pine-oak savannas. | Granite outcrops. |
Elevation | 0–150 m. (0–500 ft.) | 100–200 m. (300–700 ft.) |
Distribution |
FL; GA; NC; SC; VA
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GA |
Discussion | K. R. Park (1998) recognized both Euphorbia exserta and E. gracilior as distinct species, with the former known only from the holotype. However, this treatment follows M. J. Huft (1997) and treats E. gracilior as a synonym of E. exserta. Although the type of E. exserta is unusual in having greenish red (versus dark red) cyathia and glands, as well as small gland appendages, it is otherwise typical of the species as a whole, including features such as the upright habit, reddish coloration, scalelike proximal leaves, and filiform peduncles to 30 mm that are also common to plants formerly treated as E. gracilior. Euphorbia gracilis Elliott, which has sometimes been applied to E. exserta, is an illegitimate name (a later homonym of E. gracilis Loiseleur-Deslongchamps) and pertains here. The upright habit and usually dark red cyathia and glands distinguish Euphorbia exserta from the otherwise similar E. ipecacuanhae, while the glaucescent vegetative parts and smaller gland appendages readily separate E. exserta from the similar E. curtisii. Euphorbia exserta is found on the Gulf and Atlantic coastal plains. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia georgiana is restricted to granitic outcrops; it is known from Oglethorpe and Wilkes counties (M. H. Mayfield 2013). It is similar to E. austrotexana but has larger seeds that are much more deeply pitted and leaves that are oblanceolate instead of linear-oblanceolate to linear. It is also quite distinct from the more robust, biennial or occasionally annual E. commutata, the only other closely-related species that occurs in the area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 246. | FNA vol. 12, p. 303. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Alectoroctonum | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Tithymalopsis exserta, E. gracilior, T. gracilis | |
Name authority | (Small) Coker: Pl. Life Hartsville, 88. (1912) | Mayfield: J. Bot. Res. Inst. Texas 7: 639, fig. 2[row 2, left]. (2013) |
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