Euphorbia eriantha |
Euphorbia serpillifolia |
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beetle spurge |
thyme leaf spurge, thyme-leafed spurge, thymeleaf sandmat |
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Habit | Herbs, annual or perennial, with slender to thick, woody taproot. | Herbs, annual, with slender taproot. | ||||
Stems | erect to ascending, 10–75 cm, glabrous or with scattered appressed hairs (especially near nodes); branches arcuate. |
prostrate to ascending, often mat-forming, 7–35 cm, glabrous, pilose, or villous. |
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Leaves | alternate; petiole 0.1–0.4 mm, often indistinct, glabrous or sparsely pilose to shortly sericeous; blade linear to linear-elliptic, 20–55 × 1–3 mm, base attenuate, margins entire or with 2–4 inconspicuous teeth near apex, apex acute, abaxial surface pilose to shortly sericeous, adaxial surface usually glabrous, rarely pilose to shortly sericeous; only midvein conspicuous. |
opposite; stipules distinct, divided nearly to base into 3–5 subulate to filiform segments, these sometimes 2-fid toward apex or laciniate, 0.7–2.1 mm, glabrous; petiole 0.5–1 mm, glabrous, villous, or pilose; blade ovate, oblong, elliptic, or obovate, 3–13 × 2–7 mm, base asymmetric, rounded to oblique, margins usually entire in proximal 1/2 and serrulate in distal 1/2, rarely serrulate nearly to base, apex obtuse or truncate, surfaces often with red spot in center, not papillate, glabrous, villous, or pilose; weakly 3-veined from base, usually only midvein conspicuous. |
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Involucre | obconic, 2.1–2.6 × 1.3–2 mm, canescent; involucral lobes triangular, obscured by hairs; glands (2–)4–5, green to maroon, color often obscured by hairs, sessile and broadly attached, 0.5–0.6 × 0.5–0.6 mm, opening oblong to nearly circular, densely canescent; appendages petaloid, whitish, hoodlike, incurved and covering glands, 0.5–1 × 0.5–0.8 mm, divided into 5–12 fringed, subulate segments, densely canescent. |
obconic, 0.6–1.1 × 0.6–1 mm, glabrous, villous, or pilose; glands 4, yellow to pink, usually oblong to reniform, 0.1 × 0.2–0.3 mm; appendages white to pink, oblong or flabellate, rarely absent, (0–)0.1–0.2 × (0–)0.3–0.4 mm, distal margin entire or shallowly lobed. |
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Staminate flowers | 20–25. |
5–20. |
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Pistillate flowers | ovary canescent; styles 0.9–1.5 mm, unbranched. |
ovary glabrous, villous, or pilose; styles 0.3–0.4 mm, 2-fid 1/2 length. |
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Capsules | oblong to ovoid, 4.4–4.9 × 3.5–4.1 mm, canescent (often with interspersed glabrescent patches); columella 3.5–3.9 mm. |
broadly ovoid to oblate, cocci not elongated nor terminating in empty portion, 1.4–1.9 × 1.5–2, glabrous, pilose, or villous; columella 1.2–1.6 mm. |
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Seeds | mottled black and gray or light brown, oblong or slightly ovoid, dorsiventrally compressed in cross section, 2.8–4.1 × 2–2.4 mm, irregularly pitted and tuberculate; caruncle 0.4–0.8 × 0.6–1.1 mm. |
pink, light brown, or grayish, ovoid to narrowly ovoid, 4-angled in cross section, 1–1.4 × 0.6–0.9 mm, smooth to dimpled or rugose, or with faint transverse ridges that do not interrupt abaxial keel. |
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Cyathial | arrangement: terminal cymose branches 1, 1–2-branched; pleiochasial bracts 2–3, opposite or whorled, wholly green, similar in shape and size to distal leaves; dichasial bracts similar in shape to distal leaves or often highly reduced. |
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Cyathia | peduncle 1–1.9 mm. |
solitary or in small, cymose clusters at distal nodes or on congested, axillary branches; peduncle 0.4–1.7 mm. |
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Euphorbia eriantha |
Euphorbia serpillifolia |
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Phenology | Flowering and fruiting year-round in response to sufficient rainfall. | |||||
Habitat | Desert scrub on rocky slopes and along washes. | |||||
Elevation | 60–800 m. (200–2600 ft.) | |||||
Distribution |
AZ; CA; NM; TX; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, San Luis Potosí, Sonora)
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AZ; CA; CO; FL; GA; IA; ID; IL; MI; MN; MO; MT; ND; NE; NH; NM; NV; NY; OK; OR; PA; SD; TX; UT; WA; WI; WY; AB; BC; MB; NB; ON; QC; SK; Mexico; South America |
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Discussion | Subspecies 2 (2 in the flora). Euphorbia serpillifolia is variable, especially in regard to seed sculpturing. L. C. Wheeler (1941) documented and discussed this variation, suggesting that within the United States and Canada the taxon could be further divided into various taxa. However, Wheeler refrained from actually proposing names and commented that further study was needed. The authors concur with Wheeler and maintain a broad delimitation of the species, pending a detailed study of variation throughout its range. Otherwise indistinguishable hairy plants are treated as subsp. hirtula. In contrast to the widespread typical subspecies, subsp. hirtula ranges from northern Baja California, Mexico, to central California. In this region the two subspecies sometimes grow together. The spelling of the specific epithet follows the original publication of the name and contrasts with the often-used variant serpyllifolia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 321. | FNA vol. 12, p. 287. | ||||
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Anisophyllum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Chamaesyce serpillifolia | |||||
Name authority | Bentham: Bot. Voy. Sulphur, 51. (1844) | Persoon: Syn. Pl. 2: 14. (1806) | ||||
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