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beetle spurge

euphorbe marginée, mountain snow spurge, smoke-on-the-prairie, snow-on-the-mountain, variegated spurge, whitemargined spurge

Habit Herbs, annual or perennial, with slender to thick, woody taproot. Herbs, annual, with taproot.
Stems

erect to ascending, 10–75 cm, glabrous or with scattered appressed hairs (especially near nodes);

branches arcuate.

erect, unbranched or branched, 30–85(–150) cm, pilose or glabrous.

Leaves

alternate;

petiole 0.1–0.4 mm, often indistinct, glabrous or sparsely pilose to shortly sericeous;

blade linear to linear-elliptic, 20–55 × 1–3 mm, base attenuate, margins entire or with 2–4 inconspicuous teeth near apex, apex acute, abaxial surface pilose to shortly sericeous, adaxial surface usually glabrous, rarely pilose to shortly sericeous;

only midvein conspicuous.

alternate;

stipules 0.1–0.3 mm;

petiole 0.2–3 mm, glabrous or minutely pilose;

blade broadly ovate to elliptic, 32–62(–82) × 18–28(–52) mm, base rounded to cuneate, margins entire, often white on distal leaves, apex acute, rarely mucronate, surfaces glabrous;

venation obscure, only midvein conspicuous.

Involucre

obconic, 2.1–2.6 × 1.3–2 mm, canescent;

involucral lobes triangular, obscured by hairs;

glands (2–)4–5, green to maroon, color often obscured by hairs, sessile and broadly attached, 0.5–0.6 × 0.5–0.6 mm, opening oblong to nearly circular, densely canescent;

appendages petaloid, whitish, hoodlike, incurved and covering glands, 0.5–1 × 0.5–0.8 mm, divided into 5–12 fringed, subulate segments, densely canescent.

campanulate, 2.2–3.5 × 1.3–2.3 mm, margin between glands deeply divided into fringe of fimbriate lobes, pilose;

glands 4–5, green to greenish yellow, reniform to subcircular, 0.7–1.1 × 1–1.6 mm;

appendages white, orbiculate, 1.5–2.7 × 1.9–2.9(–3.6) mm, entire.

Staminate flowers

20–25.

30–70.

Pistillate flowers

ovary canescent;

styles 0.9–1.5 mm, unbranched.

ovary pilose;

styles 1–2.5 mm, 2-fid 1/2–2/3 length.

Capsules

oblong to ovoid, 4.4–4.9 × 3.5–4.1 mm, canescent (often with interspersed glabrescent patches);

columella 3.5–3.9 mm.

oblate, 3–5 × 3.5–7.5 mm, moderately to densely pilose;

columella 3–4.1 mm.

Seeds

mottled black and gray or light brown, oblong or slightly ovoid, dorsiventrally compressed in cross section, 2.8–4.1 × 2–2.4 mm, irregularly pitted and tuberculate;

caruncle 0.4–0.8 × 0.6–1.1 mm.

orange-tan to gray, ovoid, 3.7–3.9 × 3–3.3 mm, rugose, with 2 transverse ridges (one dark orange to brown, other inconspicuous);

caruncle absent.

Cyathial

arrangement: terminal cymose branches 1, 1–2-branched;

pleiochasial bracts 2–3, opposite or whorled, wholly green, similar in shape and size to distal leaves;

dichasial bracts similar in shape to distal leaves or often highly reduced.

Cyathia

peduncle 1–1.9 mm.

in terminal pleiochasia, dichasial bracts narrowly elliptic to oblanceolate with conspicuous white margins;

peduncle 1.8–2.7(–22) mm, densely pilose.

2n

= 56.

Euphorbia eriantha

Euphorbia marginata

Phenology Flowering and fruiting year-round in response to sufficient rainfall. Flowering and fruiting summer–fall.
Habitat Desert scrub on rocky slopes and along washes. Disturbed areas and grasslands.
Elevation 60–800 m. (200–2600 ft.) 0–1700 m. (0–5600 ft.)
Distribution
from FNA
AZ; CA; NM; TX; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, San Luis Potosí, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
AR; CA; CO; CT; IA; IL; IN; KS; LA; MA; MD; MI; MN; MO; MT; NC; NE; NH; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; UT; VA; WI; WV; WY; MB; ON; QC; SK; c Mexico; s Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia marginata is native to the central United States. The type specimen was collected by Meriwether Lewis along the Yellowstone River in southern Montana in 1806, and it has been reported to be native as far south as Arizona, New Mexico, and Texas, and as far east as southern Minnesota, western Iowa, and Missouri (G. Yatskievych 1999–2013, vol. 2). It is presumably naturalized outside of this area. Euphorbia marginata is widely cultivated as an ornamental for its showy, white-margined distal leaves, and it can escape locally.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 321. FNA vol. 12, p. 248.
Parent taxa Euphorbiaceae > Euphorbia > sect. Poinsettia Euphorbiaceae > Euphorbia > sect. Alectoroctonum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms E. bonplandii, Lepadena marginata
Name authority Bentham: Bot. Voy. Sulphur, 51. (1844) Pursh: Fl. Amer. Sept. 2: 607. (1813)
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