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beetle spurge
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royal sandmat
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| Herbs, annual or perennial, with slender to thick, woody taproot. |
Herbs, annual or short-lived perennial, with slender taproot to thickened and woody rootstock. |
erect to ascending, 10–75 cm, glabrous or with scattered appressed hairs (especially near nodes); branches arcuate. |
prostrate, usually mat-forming, terete to slightly flattened, 40 cm, lower surface glabrous, upper surface strigillose, pilose or villous. |
alternate; petiole 0.1–0.4 mm, often indistinct, glabrous or sparsely pilose to shortly sericeous; blade linear to linear-elliptic, 20–55 × 1–3 mm, base attenuate, margins entire or with 2–4 inconspicuous teeth near apex, apex acute, abaxial surface pilose to shortly sericeous, adaxial surface usually glabrous, rarely pilose to shortly sericeous; only midvein conspicuous. |
opposite; stipules distinct, entire or divided into 3–4 subulate to filiform segments, 0.8–2 mm, usually pilose, rarely glabrous; petiole 0.5–1 mm, pilose to villous; blade oblong, ovate or narrowly obovate, 3–10(–12) × 2–6 mm, base strongly asymmetric, hemicordate, margins serrulate, apex obtuse to subacute, surfaces glabrous or slightly pilose; 3-veined from base, often only midvein conspicuous. |
obconic, 2.1–2.6 × 1.3–2 mm, canescent; involucral lobes triangular, obscured by hairs; glands (2–)4–5, green to maroon, color often obscured by hairs, sessile and broadly attached, 0.5–0.6 × 0.5–0.6 mm, opening oblong to nearly circular, densely canescent; appendages petaloid, whitish, hoodlike, incurved and covering glands, 0.5–1 × 0.5–0.8 mm, divided into 5–12 fringed, subulate segments, densely canescent. |
narrowly turbinate, 1–1.2 × 0.4–0.7 mm, pilose; glands 4, yellow to pink, unequal, proximal pair oblong or linear, 0.1 × 0.3–0.4(–0.6) mm, distal pair oblong or subcircular, 0.1 × 0.1–0.2 mm; appendages pink to reddish, unequal, on proximal glands oblique, 0.4–0.8(–1) × 0.8–1.4(–2) mm, on distal glands symmetric, 0.2–0.5 × 0.2–0.5 mm, slightly undulate to slightly crenate. |
20–25. |
5–15. |
ovary canescent; styles 0.9–1.5 mm, unbranched. |
ovary pilose to strigillose in parts, glabrous in other parts; styles 0.8–1.3 mm, usually unbranched, rarely 2-fid at apex, filiform. |
oblong to ovoid, 4.4–4.9 × 3.5–4.1 mm, canescent (often with interspersed glabrescent patches); columella 3.5–3.9 mm. |
ovoid-triangular, 1.2–1.5 × 1–1.4 mm, pilose to strigillose in parts, glabrous in other parts; columella 1–1.3 mm. |
mottled black and gray or light brown, oblong or slightly ovoid, dorsiventrally compressed in cross section, 2.8–4.1 × 2–2.4 mm, irregularly pitted and tuberculate; caruncle 0.4–0.8 × 0.6–1.1 mm. |
brown to light gray, ovoid, 4-angled in cross section, 0.8–1 × 0.4–0.5 mm, with 4 or 5 deep transverse sulci alternating with low transverse ridges. |
arrangement: terminal cymose branches 1, 1–2-branched; pleiochasial bracts 2–3, opposite or whorled, wholly green, similar in shape and size to distal leaves; dichasial bracts similar in shape to distal leaves or often highly reduced. |
|
peduncle 1–1.9 mm. |
usually in small cymose clusters on congested, axillary branches; peduncle rudimentary or to 0.2 mm. |
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| Flowering and fruiting year-round in response to sufficient rainfall. |
Flowering and fruiting summer–fall. |
| Desert scrub on rocky slopes and along washes. |
Grasslands, oak forests, oak-mesquite woodlands, oak-juniper communities, rarely entering desert scrub. |
| 60–800 m. (200–2600 ft.) |
1000–2000 m. (3300–6600 ft.) |
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AZ; CA; NM; TX; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, San Luis Potosí, Sonora)
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AZ; NM; TX; Mexico
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Euphorbia indivisa is characteristic of grasslands and oak woodlands from extreme western Texas to southeastern Arizona. The species is often treated as a synonym of E. dioeca Kunth, but the two species are readily separable on the basis of their seeds. The seeds of E. indivisa possess deep transverse sulci, whereas those of E. dioeca are merely rippled or with low transverse ridges. Euphorbia dioeca is a weedy species that occurs widely throughout tropical America but has yet to be encountered within the flora area. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 12, p. 321. |
FNA vol. 12, p. 273. |
| Euphorbiaceae > Euphorbia > sect. Poinsettia |
Euphorbiaceae > Euphorbia > sect. Anisophyllum |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
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E. dioeca var. indivisa, Chamaesyce indivisa |
| Bentham: Bot. Voy. Sulphur, 51. (1844) |
(Engelmann) Tidestrom: Proc. Biol. Soc. Wash. 48: 40. (1935) |
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