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summer spurge

euphorbe réveille-matin, mad woman's milk, summer spurge, sun spurge, wart spurge, wartweed

Habit Herbs, perennial, with spreading rootstock. Herbs, annual, with taproot.
Stems

erect or ascending, unbranched, solitary or few, previous year's dead stems not persistent, 45–70 cm, usually densely puberulent to sericeous, rarely glabrous.

erect, unbranched or branched, 5–45 cm, usually glabrous or sparsely pilose.

Leaves

alternate;

stipules to 0.1 mm;

petiole (0–)1–2 mm (or absent), densely puberulent;

blade usually linear, rarely ovate, 25–55 × 1.5–4 mm, base cuneate, margins entire, revolute, apex rounded, abaxial surface glabrous or puberulent to sericeous, adaxial surface glabrous;

venation often obscure on smaller leaves, midvein conspicuous.

petiole absent or to 0.5 mm;

blade obovate-spatulate, 4–40 × 2–25 mm, base cuneate, attenuate, or auriculate, margins serrulate, apex rounded, surfaces glabrous;

venation pinnate, midvein prominent.

Involucre

campanulate, 1.2–1.4 × 1.2–2 mm, sparsely to densely puberulent;

glands 5, green, reniform, 0.2–0.3 × 0.5–0.6 mm;

appendages white, orbiculate to oblong, (0.5–)1–1.7 × 1–1.5 mm, entire.

cupulate, 1.5–2 × 0.7–1.1 mm, glabrous;

glands 4, elliptic, 0.2–0.5 × 0.5–1 mm;

horns absent.

Staminate flowers

20–25.

10–15.

Pistillate flowers

ovary glabrous or sparsely strigose;

styles 0.5–1.1 mm, 2-fid at apex to 1/2 length.

ovary glabrous;

styles 0.7–1 mm, 2-fid.

Capsules

globose, 1.8–3 × 2.5–4.8 mm, glabrous or sparsely strigose;

columella 2.3–2.5 mm.

depressed-globose, 2.5–4 × 3.2–4.2 mm, clearly 3-lobed;

cocci rounded, smooth, glabrous;

columella 0.9–1.1 mm.

Seeds

light gray, ovoid, 2 × 1.2–1.3 mm, smooth or with few, very shallow depressions;

caruncle absent.

dark brown to blackish, subovoid, 1.6–2.2 × 1.5–1.9 mm, reticulate;

caruncle elliptic, 0.9–1.1 × 0.4–0.5 mm.

Cyathia

in terminal pleiochasia;

peduncle 5–15 mm, filiform, glabrous or very sparsely puberulent to sericeous.

peduncle 0.2–1 mm.

Cyathial

arrangement: terminal pleiochasial branches (3–)5, each 1–2 times 2-branched;

pleiochasial bracts obovate, wider than distal leaves;

dichasial bracts distinct, obovate or rhombic, ± oblique, base rounded, truncate, or attenuate, margins serrulate, apex rounded;

axillary cymose branches 0.

Euphorbia discoidalis

Euphorbia helioscopia

Phenology Flowering and fruiting late spring–fall. Flowering and fruiting spring–fall.
Habitat Sand hills, pine savannas, woodland borders, open fields with sandy soils. Roadsides, waste places.
Elevation 0–150 m. (0–500 ft.) 0–1400 m. (0–4600 ft.)
Distribution
from FNA
AL; FL; GA; LA; MS; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
CA; CT; DC; DE; GA; ID; IL; LA; MA; MD; ME; MI; MT; NC; NH; NJ; NY; OH; OR; PA; SC; TN; TX; VA; VT; WA; WI; WY; AB; BC; NB; NF; NS; ON; PE; QC; SK; Europe; Asia; n Africa [Introduced in North America; introduced also in South America (Argentina, Chile)]
[WildflowerSearch map]
[BONAP county map]
Discussion

M. J. Huft (1979) remarked that Euphorbia discoidalis is uncommon west of Alabama and referred many narrow-leaved specimens from Louisiana and Texas to E. corollata. K. R. Park (1998) included them in an expanded E. discoidalis, and that is followed here. The western populations can be distinguished from E. corollata by their shorter involucral gland appendages and revolute leaf margins. Further study of these western populations is warranted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia helioscopia was collected once in Minnesota in the late 1800s but apparently did not become established there.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 245. FNA vol. 12, p. 303.
Parent taxa Euphorbiaceae > Euphorbia > sect. Alectoroctonum Euphorbiaceae > Euphorbia > subg. Esula
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Tithymalopsis discoidalis Galarhoeus helioscopius, Tithymalus helioscopius
Name authority Chapman: Fl. South. U.S., 401. (1860) Linnaeus: Sp. Pl. 1: 459. (1753)
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