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summer spurge

Garber's sandmat

Habit Herbs, perennial, with spreading rootstock. Herbs, perennial, with moderately thickened rootstock.
Stems

erect or ascending, unbranched, solitary or few, previous year's dead stems not persistent, 45–70 cm, usually densely puberulent to sericeous, rarely glabrous.

ascending, sometimes slightly woody at base, not wiry, 1–3 mm diam., 15–50 cm, canescent.

Leaves

alternate;

stipules to 0.1 mm;

petiole (0–)1–2 mm (or absent), densely puberulent;

blade usually linear, rarely ovate, 25–55 × 1.5–4 mm, base cuneate, margins entire, revolute, apex rounded, abaxial surface glabrous or puberulent to sericeous, adaxial surface glabrous;

venation often obscure on smaller leaves, midvein conspicuous.

opposite;

stipules distinct or connate at base, triangular-subulate, apex divided into 2–5 subulate to subulate-filiform segments, 0.5–0.7 mm, pilose;

petiole 0.5–1.1 mm, pilose;

blade ovate to oblong-elliptic, 4–9(–15) × 3–6 mm, base asymmetric, obtuse to rounded, margins usually entire, rarely very sparsely serrulate, apex usually obtuse to rounded, occasionally acute to apiculate, surfaces canescent; 3-veined from base, only midvein conspicuous.

Involucre

campanulate, 1.2–1.4 × 1.2–2 mm, sparsely to densely puberulent;

glands 5, green, reniform, 0.2–0.3 × 0.5–0.6 mm;

appendages white, orbiculate to oblong, (0.5–)1–1.7 × 1–1.5 mm, entire.

turbinate to campanulate, 0.6–0.9 × 0.5–0.9 mm, pilose;

glands 4, brown, slightly concave, elliptic to subcircular, 0.1–0.3 × 0.2–0.4 mm;

appendages white to pink, as narrow rim along gland, (0–)0.1–0.2 × 0.3–0.5 mm, surfaces pilose, distal margin entire or crenulate.

Staminate flowers

20–25.

8–20.

Pistillate flowers

ovary glabrous or sparsely strigose;

styles 0.5–1.1 mm, 2-fid at apex to 1/2 length.

ovary pilose;

styles 0.6–0.7 mm, 2-fid 1/2 length.

Capsules

globose, 1.8–3 × 2.5–4.8 mm, glabrous or sparsely strigose;

columella 2.3–2.5 mm.

subglobose to broadly ovoid, 1.1–1.6 × 1.3–2.1 mm, pilose;

columella 1.1–1.6 mm.

Seeds

light gray, ovoid, 2 × 1.2–1.3 mm, smooth or with few, very shallow depressions;

caruncle absent.

gray to reddish brown, oblong-ovoid, 4-angled in cross section, 1–1.2 × 0.6–0.8 mm, flat or obscurely wrinkled.

Cyathia

in terminal pleiochasia;

peduncle 5–15 mm, filiform, glabrous or very sparsely puberulent to sericeous.

solitary at nodes of short, axillary branches;

peduncle 0.2–0.4 mm.

Euphorbia discoidalis

Euphorbia garberi

Phenology Flowering and fruiting late spring–fall. Flowering and fruiting year-round.
Habitat Sand hills, pine savannas, woodland borders, open fields with sandy soils. Beach dunes, coastal rock barrens, disturbed uplands and pine rocklands.
Elevation 0–150 m. (0–500 ft.) 0–10 m. (0–0 ft.)
Distribution
from FNA
AL; FL; GA; LA; MS; TX
[WildflowerSearch map]
[BONAP county map]
from FNA
FL
[BONAP county map]
Discussion

M. J. Huft (1979) remarked that Euphorbia discoidalis is uncommon west of Alabama and referred many narrow-leaved specimens from Louisiana and Texas to E. corollata. K. R. Park (1998) included them in an expanded E. discoidalis, and that is followed here. The western populations can be distinguished from E. corollata by their shorter involucral gland appendages and revolute leaf margins. Further study of these western populations is warranted.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia garberi appears to be an interspecific hybrid, but its parentage is not entirely clear (Y. Yang and P. E. Berry 2011). The taxon is closely related to E. blodgettii, E. porteriana, and E. serpens. It is restricted to Collier, Miami-Dade, and Monroe counties in southern Florida and is federally listed as threatened due to impacts from habitat loss, fire suppression, and invasive species. A. Herndon (1993) synonymized E. porteriana var. keyensis under E. garberi because of its uniformly pilose capsules, and that treatment is followed here.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 245. FNA vol. 12, p. 268.
Parent taxa Euphorbiaceae > Euphorbia > sect. Alectoroctonum Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Tithymalopsis discoidalis Chamaesyce adicioides, C. brachypoda, C. garberi, C. keyensis, C. mosieri, C. porteriana var. keyensis, E. porteriana var. keyensis
Name authority Chapman: Fl. South. U.S., 401. (1860) Engelmann ex Chapman: Fl. South. U.S. ed. 2, 646. (1883)
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