Euphorbia dentata |
Euphorbia spathulata |
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green poinsettia, tooth poinsettia, tooth spurge |
prairie spurge, reticulate seed spurge, spatulate leaf spurge, warty spurge |
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Habit | Herbs, annual, with taproot. | Herbs, usually annual, rarely biennial, with taproot. |
Stems | erect or ascending, 15–60 cm, both pilose and inconspicuously strigillose; branches usually ± straight, occasionally proximal branches arcuate. |
erect or ascending, unbranched or branched, 10–70 cm, glabrous. |
Leaves | usually opposite, occasionally alternate at distal nodes; petiole 5–20 mm, pilose; blade 30–70 × 4–35 mm, narrowly lanceolate to suborbiculate, usually broadest below middle, base usually acute to subobtuse, rarely subtruncate, margins coarsely crenate-dentate or doubly crenate, strigillose, flat to slightly revolute, apex broadly acute, abaxial surface long pilose with weak, filiform hairs, adaxial surface sparsely pilose to glabrate; venation pinnate, midvein prominent. |
petiole absent or to 0.2 mm; blade oblanceolate, oblong-oblanceolate, spatulate, or cuneate, 10–50 × 6–11 mm, base broadly attenuate to rounded or shallowly cordate-clasping, margins finely serrulate (usually distally), apex usually rounded to obtuse, occasionally slightly retuse or obcordate proximally, bluntly mucronate, surfaces glabrous; venation pinnate, midvein prominent. |
Involucre | campanulate, 3.8 × 1.8 mm, glabrous; involucral lobes divided into several linear, smooth lobes; glands (1–)2, green, sessile and broadly attached, 0.7–0.9 × 0.9–1.2 mm, opening oblong, glabrous; appendages absent. |
campanulate to cupulate, 0.6–1(–1.5) × 0.8–1.2 mm, glabrous; glands 4–5, elliptic, oblong, to slightly reniform, 0.2–0.6 × 0.4–1 mm; horns absent. |
Staminate flowers | 8–10. |
3–10. |
Pistillate flowers | ovary glabrous, styles 1.2 mm, 2-fid 1/2 to nearly entire length. |
ovary glabrous; styles 0.8–1.5 mm, 2-fid. |
Capsules | depressed-globose, 2.5–2.8 × 3.5–4 mm, 3-lobed, glabrous; columella 1.8–2.1 mm. |
depressed-globose, 2–3.5 × 4 mm, 3-lobed; cocci rounded, verrucose, verrucae 0.1–0.2 mm, glabrous; columella 1.4–2.2 mm. |
Seeds | pale gray to black, ovoid, rounded in cross section, 2.1–2.7 × 1.7–2.1 mm, evenly minute-tuberculate; caruncle 0.4–0.6 mm. |
red-brown to dark purple, occasionally ± glaucous, broadly ellipsoid-ovoid to nearly globose, 1.3–2.5 × 1.5–1.8 mm, smooth, reticulate, or finely low-ridged; caruncle irregularly reniform to round, subconic to lenticular, 0.3–0.4 × 0.5–0.6 mm. |
Cyathial | arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green, white, or mauve at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts similar in shape to distal leaves but smaller (often highly reduced). |
arrangement: terminal pleiochasial branches 3(–5), each 1–3 times 2-branched; pleiochasial bracts broadly ovate to ovate-oblong, shorter and wider than distal leaves; dichasial bracts distinct, broadly ovate, ovate-triangular, or ovate-elliptic, base cordate-clasping or subcordate to rounded, margins serrulate, apex rounded to obtuse or acute; axillary cymose branches (0–4)5–12. |
Cyathia | peduncle 0.7–1 mm. |
peduncle 0.3–1(–1.5) mm. |
2n | = 28. |
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Euphorbia dentata |
Euphorbia spathulata |
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Phenology | Flowering and fruiting spring–fall. | Flowering and fruiting spring–summer. |
Habitat | Bottomland forests, stream and river banks, bluffs, prairies, glades, fallow fields, roadsides, railroad cinders, open disturbed areas. | Forests, fallow fields, prairies, pastures, glades, stream banks, waste places, roadsides. |
Elevation | 0–1000 m. (0–3300 ft.) | 0–3500 m. (0–11500 ft.) |
Distribution |
AL; AR; GA; IA; IL; IN; KS; KY; LA; MD; MI; MO; MS; NC; NE; OH; OK; PA; SC; TN; TX; VA; WV; ON; Mexico (Chihuahua, Coahuila, Nuevo León, Tamaulipas)
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AL; AR; AZ; CA; CO; DC; FL; GA; IA; ID; IL; IN; KS; KY; LA; MD; MI; MN; MO; MS; MT; NC; ND; NE; NM; OH; OK; OR; PA; SC; SD; TN; TX; UT; VA; WA; WI; WV; WY; ON; Mexico (Chihuahua, Sonora); s South America
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Discussion | Euphorbia dentata is native from northern Mexico and the south central United States north and east through the Ohio River Valley. Scattered occurrences in the southeastern United States likely represent adventive populations. Reports of E. dentata as a noxious weed (from the United States and the Old World) should most likely be attributed to introductions of E. davidii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
As treated here, Euphorbia spathulata is a wide-ranging and variable species. J. B. S. Norton (1900) recognized a number of segregates (for example, E. arkansana, E. dictyosperma, and E. obtusata) all of which are included here in a broadly defined E. spathulata. The only segregate species that has been widely recognized in regional floras is E. obtusata (for example, M. L. Fernald 1950; T. S. Cooperrider 1995; G. Yatskievych 1999–2013, vol. 2). Authors have generally distinguished the eastern North American E. obtusata from the western E. spathulata by the former's larger seeds (1.7–2.3 mm versus 1.5–1.7 mm) with smooth (versus reticulate) surfaces, larger involucres, red (versus yellow) involucral glands, and cordate-clasping (versus rounded to subcordate) dichasial bracts. Examination of specimens of E. spathulata in the broad sense from throughout North America showed that there is some geographic patterning to seed size and surface sculpturing, but the variation does not segregate cleanly into two discrete taxa. Plants from western North America typically have small seeds (1.5–1.7 mm) with reticulate surfaces, although some western individuals have seeds 1.8–1.9 mm long with reticulate surfaces. Plants from Texas generally have small seeds (1.5–1.6 mm) but with the surfaces either reticulate or completely smooth. Plants from adjacent Louisiana have small seeds with faintly reticulate to almost bumpy surfaces. Plants from eastern North America have larger seeds (2–2.3 mm) with usually smooth surfaces, although individuals from Tennessee and the Carolinas have faintly reticulate surfaces. Involucre height, gland color, and the shape of the dichasial bracts do not segregate with seed size as previous treatments have suggested. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 321. | FNA vol. 12, p. 309. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Poinsettia dentata | E. arkansana, E. dictyosperma, E. obtusata, Galarhoeus arkansanus, G. obtusatus, Tithymalus arkansanus, T. dictyospermus, T. obtusatus, T. spathulatus |
Name authority | Michaux: Fl. Bor.-Amer. 2: 211. (1803) | Lamarck: in J. Lamarck et al., Encycl. 2: 428. (1788) |
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