Euphorbia dentata |
Euphorbia exstipulata |
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green poinsettia, tooth poinsettia, tooth spurge |
square-seed spurge, squareseed or Clark Mountain spurge |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with slender taproot. |
Stems | erect or ascending, 15–60 cm, both pilose and inconspicuously strigillose; branches usually ± straight, occasionally proximal branches arcuate. |
erect, 5–26 cm, uniformly puberulent to hispidulous or glabrous; branches arcuate. |
Leaves | usually opposite, occasionally alternate at distal nodes; petiole 5–20 mm, pilose; blade 30–70 × 4–35 mm, narrowly lanceolate to suborbiculate, usually broadest below middle, base usually acute to subobtuse, rarely subtruncate, margins coarsely crenate-dentate or doubly crenate, strigillose, flat to slightly revolute, apex broadly acute, abaxial surface long pilose with weak, filiform hairs, adaxial surface sparsely pilose to glabrate; venation pinnate, midvein prominent. |
opposite; petiole 1–3 mm, often indistinct, glabrous or puberulent; blade linear to narrowly elliptic or ovate, 14–42 × 3–28 mm, base attenuate, margins coarsely serrate, occasionally revolute, apex acute to obtuse, abaxial surface sparsely hispidulous to strigillose, adaxial surface glabrous; midvein conspicuous. |
Involucre | campanulate, 3.8 × 1.8 mm, glabrous; involucral lobes divided into several linear, smooth lobes; glands (1–)2, green, sessile and broadly attached, 0.7–0.9 × 0.9–1.2 mm, opening oblong, glabrous; appendages absent. |
turbinate to campanulate, 1.1–1.5 × 1–1.3 mm, glabrous, pilose or puberulent; involucral lobes divided into several linear lobes; glands 4(–5), yellow to pink, sessile and broadly attached, 0.2 × 0.3–0.4 mm, opening oblong to nearly circular, glabrous; appendages usually petaloid, white to pink, ovate to trapezoidal, occasionally absent, not incurved and covering glands, 0.2–0.4 × 0.3–0.8 mm, entire, undulate, or conspicuously divided into triangular segments, glabrous. |
Staminate flowers | 8–10. |
10–12. |
Pistillate flowers | ovary glabrous, styles 1.2 mm, 2-fid 1/2 to nearly entire length. |
ovary puberulent on keels, styles 0.8–1.1 mm, 2-fid 1/2 to nearly entire length. |
Capsules | depressed-globose, 2.5–2.8 × 3.5–4 mm, 3-lobed, glabrous; columella 1.8–2.1 mm. |
broadly depressed-oblong to ovoid, 2.7–3.3 × 3.1–3.9 mm, puberulent (with appressed hairs usually concentrated on keels); columella 1.9–2.5 mm. |
Seeds | pale gray to black, ovoid, rounded in cross section, 2.1–2.7 × 1.7–2.1 mm, evenly minute-tuberculate; caruncle 0.4–0.6 mm. |
white to gray or light brown, ovoid, bluntly 4-angled in cross section, 1.9–2.5 × 1.4–1.7 mm, tuberculate, often with 2 transverse ridges; caruncle 0.1 × 0.2 mm. |
Cyathial | arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green, white, or mauve at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts similar in shape to distal leaves but smaller (often highly reduced). |
arrangement: terminal cymose or dichasial branches usually 1–2, occasionally reduced to monochasia, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts similar in shape to distal leaves but smaller or highly reduced. |
Cyathia | peduncle 0.7–1 mm. |
peduncle 1–1.9 mm. |
2n | = 28. |
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Euphorbia dentata |
Euphorbia exstipulata |
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Phenology | Flowering and fruiting spring–fall. | Flowering and fruiting summer–fall. |
Habitat | Bottomland forests, stream and river banks, bluffs, prairies, glades, fallow fields, roadsides, railroad cinders, open disturbed areas. | Desert scrub, grasslands, mesquite savannas, oak and oak-juniper woodlands. |
Elevation | 0–1000 m. (0–3300 ft.) | 800–2000 m. (2600–6600 ft.) |
Distribution |
AL; AR; GA; IA; IL; IN; KS; KY; LA; MD; MI; MO; MS; NC; NE; OH; OK; PA; SC; TN; TX; VA; WV; ON; Mexico (Chihuahua, Coahuila, Nuevo León, Tamaulipas)
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AZ; CA; NM; TX; UT; WY; Mexico (Chihuahua, Coahuila, Durango, Guanajuato, San Luis Potosí, Sonora, Zacatecas)
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Discussion | Euphorbia dentata is native from northern Mexico and the south central United States north and east through the Ohio River Valley. Scattered occurrences in the southeastern United States likely represent adventive populations. Reports of E. dentata as a noxious weed (from the United States and the Old World) should most likely be attributed to introductions of E. davidii. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia exstipulata is native from Texas to California and northern Mexico. The species was found once in the late nineteenth century in Wyoming but has not been re-collected there. Broad-leaved plants have been segregated as var. lata, but the variation in leaf shape is continuous and no varieties are formally recognized here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 321. | FNA vol. 12, p. 322. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Poinsettia |
Sibling taxa | ||
Synonyms | Poinsettia dentata | E. exstipulata var. lata |
Name authority | Michaux: Fl. Bor.-Amer. 2: 211. (1803) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) |
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