Euphorbia davidii |
Euphorbia yaquiana |
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David's poinsettia, David's spurge, tooth poinsettia, tooth spurge |
hairy Mojave spurge |
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Habit | Herbs, annual, with taproot. | Herbs, perennial, with thick rootstock. |
Stems | erect or ascending, 20–70 cm, both coarsely and sparsely hirsute and closely strigillose; branches usually ± straight, occasionally proximal branches arcuate. |
slender, erect or ascending, sometimes sinuous, densely branched near base, 10–50 cm, moderately to densely puberulent to lanulose. |
Leaves | usually opposite, occasionally alternate at distal nodes; petiole 7–25 mm, strigose; blade usually narrowly to broadly elliptic, occasionally lance-elliptic, 10–100 × 5–35 mm, base cuneate to attenuate, margins coarsely crenate-dentate, strigose, revolute to nearly flat, apex broadly acute to acuminate, or obtuse, abaxial surface strigose with stiff, strongly tapered hairs, adaxial surface sparsely strigose-hirsute; venation pinnate, midvein prominent. |
petiole 0–1 mm; blade usually lanceolate or elliptic-lanceolate, sometimes slightly oblanceolate, 8–30 × 6–14 mm, base usually acute, occasionally short-attenuate, rarely obtuse, margins entire, apex usually acute, occasionally obtuse, acuminate, or cuspidate, surfaces sparsely to moderately puberulent to lanulose; venation pinnate, sometimes obscure, midvein prominent. |
Involucre | cylindric, 2.5–3 × 1.3–1.8 mm, glabrous; involucral lobes divided into 5–7 linear, papillate lobes; gland 1, yellow-green, sessile and broadly attached, 0.9 × 1.3 mm, opening oblong, glabrous; appendages absent. |
campanulate to broadly turbinate, 2.2–3 × 2–2.5 mm, puberulent to lanulose; glands 4, semicircular, trapezoidal, or elliptic-truncate, 0.8–1.5 × 1–2.2 mm, margins strongly crenate or dentate; horns usually absent, if present then straight, 0.1–0.2 mm, generally equaling teeth on gland margin. |
Staminate flowers | 5–8. |
12–20. |
Pistillate flowers | ovary glabrous or sparsely strigose; styles 1 mm, 2-fid 1/2 to nearly entire length. |
ovary usually puberulent, occasionally lanulose; styles 1–1.2 mm, 2-fid. |
Capsules | broadly ovoid, 2.9–3.3 × 4–4.8 mm, 3-lobed, glabrous; columella 2.2–2.7 mm. |
oblong-ovoid, 3.5–4 × 3–4 mm, 3-lobed; cocci rounded, smooth, usually puberulent, occasionally lanulose; columella 2.5–3 mm. |
Seeds | black to brown or pale gray, ovoid to triangular-ovoid, angular in cross section, 2.4–2.9 × 2.2–2.9 mm, low-tuberculate, tubercles irregularly arranged or in faint, transverse row; caruncle 0.9–1.1 mm. |
gray to whitish, oblong cylindric, 2–3 × 1.5–1.8 mm, irregularly shallowly pitted to almost smooth; caruncle conic, 0.6 × 0.6 mm. |
Cyathial | arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched; pleiochasial bracts 2–4, often whorled, green with diffuse greenish white to mauve near base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts similar in shape to distal leaves but smaller, often highly reduced. |
arrangement: terminal pleiochasial branches 3–5, each 1–2 times 2-branched; pleiochasial bracts broadly ovate to subcordate, usually similar in size to, occasionally wider than, distal leaves; dichasial bracts distinct, broadly ovate to almost reniform, base obtuse, margins entire, apex obtuse, acuminate to cuspidate; axillary cymose branches 0–5. |
Cyathia | peduncle 0.5–1 mm. |
peduncle 0.3–0.8 mm. |
2n | = 56. |
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Euphorbia davidii |
Euphorbia yaquiana |
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Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting spring–summer. |
Habitat | Forests, stream and riverbanks, prairies, roadsides and open disturbed areas. | Ponderosa pine forests, oak-pine mixed forests, dry stream banks and beds, open scrub areas, roadsides. |
Elevation | 200–1500 m. (700–4900 ft.) | 1000–2200 m. (3300–7200 ft.) |
Distribution |
AR; AZ; CA; CO; FL; IA; ID; IL; IN; KS; KY; LA; MA; MI; MN; MO; NC; NE; NJ; NM; NY; OH; OK; SD; TN; TX; UT; VA; VT; WI; WV; WY; ON; QC; Mexico (Chihuahua, Coahuila, Sonora) [Introduced in South America, Eurasia (China, Russia), Australia]
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AZ |
Discussion | Euphorbia davidii is native from the southwestern United States and northern Mexico north through the southern Great Plains; it apparently is adventive elsewhere. The species is the weediest member of the E. dentata species group (following M. H. Mayfield 1997) and has become an agricultural weed in North America, South America (for example, Argentina), and in the Old World (particularly Australia and Russia). Euphorbia davidii can be distinguished from the closely similar E. dentata by its larger capsules and seeds, often more elliptic leaves, and shorter, stiffer hairs. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia yaquiana is endemic to Pima and Graham counties in southern Arizona and is known only from the Santa Catalina and Pinaleño mountains. Records of E. yaquiana from southwestern Colorado (as E. incisa var. mollis) likely represent misidentifications of E. brachycera; therefore, those disjunct occurrences have been excluded here from the distribution of E. yaquiana. Euphorbia yaquiana has often been treated as a synonym of E. schizoloba var. mollis, but molecular phylogenetic data show that it is more closely related to E. brachycera and E. chamaesula (J. A. Peirson et al. 2014). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 320. | FNA vol. 12, p. 313. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | E. schizoloba var. mollis, E. incisa var. mollis | |
Name authority | Subils: Kurtziana 17: 125, figs. 1, 2H–J. (1984) | Tidestrom: Proc. Biol. Soc. Wash. 48: 41. (1935) |
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