Euphorbia davidii |
Euphorbia theriaca |
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David's poinsettia, David's spurge, tooth poinsettia, tooth spurge |
Terlingua sandmat, Terlingua spurge |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with slender taproot. | ||||
Stems | erect or ascending, 20–70 cm, both coarsely and sparsely hirsute and closely strigillose; branches usually ± straight, occasionally proximal branches arcuate. |
prostrate to reclining, not mat-forming, 5–30 cm, glabrous. |
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Leaves | usually opposite, occasionally alternate at distal nodes; petiole 7–25 mm, strigose; blade usually narrowly to broadly elliptic, occasionally lance-elliptic, 10–100 × 5–35 mm, base cuneate to attenuate, margins coarsely crenate-dentate, strigose, revolute to nearly flat, apex broadly acute to acuminate, or obtuse, abaxial surface strigose with stiff, strongly tapered hairs, adaxial surface sparsely strigose-hirsute; venation pinnate, midvein prominent. |
opposite; stipules usually distinct, occasionally connate basally on lower side of stem, subulate or scalelike, usually entire, occasionally 2-fid or margin sparsely ciliate, 0.4–1 mm, glabrous; petiole 0.7–1.2(–1.5) mm, glabrous; blade ovate, oblong, orbiculate, or obovate, 2–7.1 × 1–3.5 mm, base slightly asymmetric, rounded, margins entire, often revolute on drying, apex usually rounded, occasionally slightly emarginate, surfaces glabrous; venation usually obscure, only midvein conspicuous. |
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Involucre | cylindric, 2.5–3 × 1.3–1.8 mm, glabrous; involucral lobes divided into 5–7 linear, papillate lobes; gland 1, yellow-green, sessile and broadly attached, 0.9 × 1.3 mm, opening oblong, glabrous; appendages absent. |
usually turbinate-campanulate to hemispheric, occasionally suburceolate, 1–1.8 × 0.9–1.4 mm, glabrous; glands 4, yellow-green to red-purple, sessile or short-stipitate, subcircular to slightly elliptic, 0.2–0.5 × 0.2–0.7 mm; appendages absent or white to pink, semilunate or forming rim at edge of gland, (0–)0.1–0.4 × (0–)0.3–0.9 mm, entire or slightly crenate. |
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Staminate flowers | 5–8. |
15–36. |
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Pistillate flowers | ovary glabrous or sparsely strigose; styles 1 mm, 2-fid 1/2 to nearly entire length. |
ovary glabrous; styles 0.3–0.5 mm, 2-fid 1/2 length. |
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Capsules | broadly ovoid, 2.9–3.3 × 4–4.8 mm, 3-lobed, glabrous; columella 2.2–2.7 mm. |
broadly ovoid, 1.1–1.6 × 1.5–1.8 mm, glabrous; columella 1.2–1.5 mm. |
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Seeds | black to brown or pale gray, ovoid to triangular-ovoid, angular in cross section, 2.4–2.9 × 2.2–2.9 mm, low-tuberculate, tubercles irregularly arranged or in faint, transverse row; caruncle 0.9–1.1 mm. |
whitish, reddish brown beneath coat, ovate, 4-angled in cross section, 0.8–1.2 × 0.5–0.8 mm, with (2–)3(–5) deep transverse ridges. |
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Cyathial | arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched; pleiochasial bracts 2–4, often whorled, green with diffuse greenish white to mauve near base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts similar in shape to distal leaves but smaller, often highly reduced. |
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Cyathia | peduncle 0.5–1 mm. |
usually solitary at distal nodes, rarely clustered on short, axillary branches; peduncle 0.3–1.3 mm. |
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2n | = 56. |
= 28. |
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Euphorbia davidii |
Euphorbia theriaca |
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Phenology | Flowering and fruiting summer–fall. | |||||
Habitat | Forests, stream and riverbanks, prairies, roadsides and open disturbed areas. | |||||
Elevation | 200–1500 m. (700–4900 ft.) | |||||
Distribution |
AR; AZ; CA; CO; FL; IA; ID; IL; IN; KS; KY; LA; MA; MI; MN; MO; NC; NE; NJ; NM; NY; OH; OK; SD; TN; TX; UT; VA; VT; WI; WV; WY; ON; QC; Mexico (Chihuahua, Coahuila, Sonora) [Introduced in South America, Eurasia (China, Russia), Australia]
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NM; TX; n Mexico
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Discussion | Euphorbia davidii is native from the southwestern United States and northern Mexico north through the southern Great Plains; it apparently is adventive elsewhere. The species is the weediest member of the E. dentata species group (following M. H. Mayfield 1997) and has become an agricultural weed in North America, South America (for example, Argentina), and in the Old World (particularly Australia and Russia). Euphorbia davidii can be distinguished from the closely similar E. dentata by its larger capsules and seeds, often more elliptic leaves, and shorter, stiffer hairs. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 320. | FNA vol. 12, p. 290. | ||||
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Anisophyllum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Chamaesyce theriaca | |||||
Name authority | Subils: Kurtziana 17: 125, figs. 1, 2H–J. (1984) | L. C. Wheeler: Rhodora 43: 242, plate 660, fig. A. (1941) | ||||
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