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David's poinsettia, David's spurge, tooth poinsettia, tooth spurge

beetle spurge

Habit Herbs, annual, with taproot. Herbs, annual or perennial, with slender to thick, woody taproot.
Stems

erect or ascending, 20–70 cm, both coarsely and sparsely hirsute and closely strigillose;

branches usually ± straight, occasionally proximal branches arcuate.

erect to ascending, 10–75 cm, glabrous or with scattered appressed hairs (especially near nodes);

branches arcuate.

Leaves

usually opposite, occasionally alternate at distal nodes;

petiole 7–25 mm, strigose;

blade usually narrowly to broadly elliptic, occasionally lance-elliptic, 10–100 × 5–35 mm, base cuneate to attenuate, margins coarsely crenate-dentate, strigose, revolute to nearly flat, apex broadly acute to acuminate, or obtuse, abaxial surface strigose with stiff, strongly tapered hairs, adaxial surface sparsely strigose-hirsute;

venation pinnate, midvein prominent.

alternate;

petiole 0.1–0.4 mm, often indistinct, glabrous or sparsely pilose to shortly sericeous;

blade linear to linear-elliptic, 20–55 × 1–3 mm, base attenuate, margins entire or with 2–4 inconspicuous teeth near apex, apex acute, abaxial surface pilose to shortly sericeous, adaxial surface usually glabrous, rarely pilose to shortly sericeous;

only midvein conspicuous.

Involucre

cylindric, 2.5–3 × 1.3–1.8 mm, glabrous;

involucral lobes divided into 5–7 linear, papillate lobes;

gland 1, yellow-green, sessile and broadly attached, 0.9 × 1.3 mm, opening oblong, glabrous;

appendages absent.

obconic, 2.1–2.6 × 1.3–2 mm, canescent;

involucral lobes triangular, obscured by hairs;

glands (2–)4–5, green to maroon, color often obscured by hairs, sessile and broadly attached, 0.5–0.6 × 0.5–0.6 mm, opening oblong to nearly circular, densely canescent;

appendages petaloid, whitish, hoodlike, incurved and covering glands, 0.5–1 × 0.5–0.8 mm, divided into 5–12 fringed, subulate segments, densely canescent.

Staminate flowers

5–8.

20–25.

Pistillate flowers

ovary glabrous or sparsely strigose;

styles 1 mm, 2-fid 1/2 to nearly entire length.

ovary canescent;

styles 0.9–1.5 mm, unbranched.

Capsules

broadly ovoid, 2.9–3.3 × 4–4.8 mm, 3-lobed, glabrous;

columella 2.2–2.7 mm.

oblong to ovoid, 4.4–4.9 × 3.5–4.1 mm, canescent (often with interspersed glabrescent patches);

columella 3.5–3.9 mm.

Seeds

black to brown or pale gray, ovoid to triangular-ovoid, angular in cross section, 2.4–2.9 × 2.2–2.9 mm, low-tuberculate, tubercles irregularly arranged or in faint, transverse row;

caruncle 0.9–1.1 mm.

mottled black and gray or light brown, oblong or slightly ovoid, dorsiventrally compressed in cross section, 2.8–4.1 × 2–2.4 mm, irregularly pitted and tuberculate;

caruncle 0.4–0.8 × 0.6–1.1 mm.

Cyathial

arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched;

pleiochasial bracts 2–4, often whorled, green with diffuse greenish white to mauve near base, similar in shape and size to distal leaves or slightly narrower;

dichasial bracts similar in shape to distal leaves but smaller, often highly reduced.

arrangement: terminal cymose branches 1, 1–2-branched;

pleiochasial bracts 2–3, opposite or whorled, wholly green, similar in shape and size to distal leaves;

dichasial bracts similar in shape to distal leaves or often highly reduced.

Cyathia

peduncle 0.5–1 mm.

peduncle 1–1.9 mm.

2n

= 56.

Euphorbia davidii

Euphorbia eriantha

Phenology Flowering and fruiting summer–fall. Flowering and fruiting year-round in response to sufficient rainfall.
Habitat Forests, stream and riverbanks, prairies, roadsides and open disturbed areas. Desert scrub on rocky slopes and along washes.
Elevation 200–1500 m. (700–4900 ft.) 60–800 m. (200–2600 ft.)
Distribution
from FNA
AR; AZ; CA; CO; FL; IA; ID; IL; IN; KS; KY; LA; MA; MI; MN; MO; NC; NE; NJ; NM; NY; OH; OK; SD; TN; TX; UT; VA; VT; WI; WV; WY; ON; QC; Mexico (Chihuahua, Coahuila, Sonora) [Introduced in South America, Eurasia (China, Russia), Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NM; TX; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, San Luis Potosí, Sonora)
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia davidii is native from the southwestern United States and northern Mexico north through the southern Great Plains; it apparently is adventive elsewhere. The species is the weediest member of the E. dentata species group (following M. H. Mayfield 1997) and has become an agricultural weed in North America, South America (for example, Argentina), and in the Old World (particularly Australia and Russia). Euphorbia davidii can be distinguished from the closely similar E. dentata by its larger capsules and seeds, often more elliptic leaves, and shorter, stiffer hairs.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 320. FNA vol. 12, p. 321.
Parent taxa Euphorbiaceae > Euphorbia > sect. Poinsettia Euphorbiaceae > Euphorbia > sect. Poinsettia
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Name authority Subils: Kurtziana 17: 125, figs. 1, 2H–J. (1984) Bentham: Bot. Voy. Sulphur, 51. (1844)
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