Euphorbia cyathophora |
Euphorbia pubentissima |
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fire on the mountain, painted leaf, painted poinsettia |
false flowering spurge, southeastern flowering spurge |
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Habit | Herbs, annual, with spreading taproots. | Herbs, perennial, with spreading rootstock. |
Stems | erect or ascending, 20–100 cm, glabrous, sparsely pilose, or puberulent; branches ± straight. |
erect, usually unbranched, occasionally few branched distally, solitary or few, previous year's dead stems not persistent, 30–65 cm, usually glabrous, rarely villous. |
Leaves | usually alternate, occasionally opposite proximally; petiole 2–20 mm, glabrous or pilose, or often hispid abaxially near blade junction; blade linear, lanceolate, elliptic, or wider leaves pandurate and unequally 4-lobed, occasionally polymorphic on single plants, 15–250 × 4–40 mm, base acute to cuneate, margins subulately glandular-serrulate distally, or sparsely glandular and subentire, hirtellous to glabrate, flat to revolute, apex acute to cuneate, abaxial surface sparsely pilose or glabrate, adaxial surface glabrous or sparsely puberulent; venation pinnate, midvein prominent. |
alternate; stipules 1 mm; petiole (0–)1–2(–10) mm, glabrous or densely villous; blade lanceolate, lance-ovate, elliptic, or obovate, 40–68 × 6–18 mm, often reflexed, base cuneate to rounded, margins entire, apex usually rounded, sometimes broadly acute, abaxial surface glabrous or villous, adaxial surface usually glabrous, rarely sparsely villous; venation occasionally obscure on small leaves, midvein conspicuous. |
Involucre | campanulate, occasionally broadly so, 1.8–2.8 × 2.2–2.8 mm, glabrous; involucral lobes triangularly 3–5 lobed; gland 1, yellow-green, sessile to substipitate and narrowly to broadly attached, 1–1.4 × 0.9–1.6 mm, opening oblong (flattened without pressing), without annular rim, glabrous; appendages absent. |
campanulate or hemispheric, 1.3–1.7 × 1.8–2.2 mm, glabrous or villous; glands 5 (7–10 on central cyathium), green, reniform or broadly elliptic, 0.3–0.5 × 0.5–0.8 mm; appendages white, orbiculate or narrowly flabellate, 1–2.2 × 1.5 mm, entire. |
Staminate flowers | 7–20. |
20–25. |
Pistillate flowers | ovary glabrous; styles 1.6 mm, 2-fid nearly entire length. |
ovary glabrous or sparsely villous; styles 0.6–1.1 mm, 2-fid 1/2 length. |
Capsules | green, depressed-globose to ellipsoid, 2.8–3.2 × 4–4.5 mm, 3-lobed, glabrous; columella 2–2.7 mm. |
globose, 2–2.4 × 3.3–4.8 mm, glabrous or sparsely villous; columella 1.8–2.1 mm. |
Seeds | black to ashy gray or light brown, cylindric to ovoid, rounded in cross section, 2.3–3.1 × 1.9–2.5 mm, uniformly tuberculate or tubercles arranged in median, transverse ridge in cylindric seeds; caruncle absent. |
light gray, ovoid, 2.2–2.4 × 1.6–1.8 mm, with shallow depressions; caruncle absent. |
Cyathial | arrangement: terminal pleiochasial branches (1–)3, 1–2-branched; pleiochasial bracts 2–3(–4), often as tight, involucrate whorl, usually green with white, pink, or red at base, occasionally distal bracts wholly white, pink, or red, rarely all bracts wholly green, similar in shape and size to distal leaves; dichasial bracts often colored, similar in shape and size to distal stem leaves or highly reduced. |
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Cyathia | peduncle 1.6–2.8 mm. |
in terminal pleiochasia; peduncle 1–5 mm (or 15–40 mm on early flowering individuals), usually glabrous, occasionally sparsely villous. |
2n | = 28, 56. |
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Euphorbia cyathophora |
Euphorbia pubentissima |
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Phenology | Flowering and fruiting late spring–fall. | Flowering and fruiting spring–fall. |
Habitat | Bottomland forests, stream and river banks, bases of bluffs, fallow fields, roadsides, open disturbed areas. | Open fields, cliffs, woods, floodplains. |
Elevation | 0–1800 m. (0–5900 ft.) | 0–900 m. (0–3000 ft.) |
Distribution |
AL; AR; CA; FL; GA; IA; IL; IN; KS; KY; LA; MD; MN; MO; MS; NC; NE; NM; OH; OK; SC; SD; TN; TX; UT; VA; WI; Mexico; Central America; South America; West Indies [Introduced in Eurasia, Africa, Indian Ocean Islands, Pacific Islands, Australia]
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AL; FL; GA; LA; MD; MS; NC; SC; TN; VA; WV
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Discussion | Euphorbia cyathophora is native to the midwestern and southeastern United States, Mexico, the West Indies, and northern South America. Leaf shape can be polymorphic on individuals of this species, but not to the extent as in E. heterophylla. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The taxonomic history of Euphorbia pubentissima is complex. The species has been included within a very broadly defined E. corollata in the past. Therefore, its geographic distribution appears significantly more wide-ranging in some treatments. The framework established by M. J. Huft (1979) and later by K. R. Park (1998) is followed here. Euphorbia pubentissima is recognized as a variable species restricted to the southeastern United States. The species can be distinguished from E. corollata by its shorter involucral gland appendages and smaller seeds. Euphorbia paniculata Elliott, which sometimes is applied to E. pubentissima, is an illegitimate name (a later homonym of Euphorbia paniculata Desfontaines). (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 319. | FNA vol. 12, p. 250. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Synonyms | Poinsettia cyathophora | E. apocynifolia, E. corollata var. paniculata, E. corollata var. zinniiflora, E. zinniiflora, Tithymalopsis apocynifolia, T. paniculata, T. zinniiflora |
Name authority | Murray: Commentat. Soc. Regiae Sci. Gott. 7: 81, plate 1. (1786) | Michaux: Fl. Bor.-Amer. 2: 212. (1803) |
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