Euphorbia cyathophora |
Euphorbia lurida |
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fire on the mountain, painted leaf, painted poinsettia |
woodland spurge |
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Habit | Herbs, annual, with spreading taproots. | Herbs, perennial, with thick rootstock. |
Stems | erect or ascending, 20–100 cm, glabrous, sparsely pilose, or puberulent; branches ± straight. |
erect or ascending, unbranched, sometimes sinuous, 5–30 cm, glabrous or sparsely to densely puberulent. |
Leaves | usually alternate, occasionally opposite proximally; petiole 2–20 mm, glabrous or pilose, or often hispid abaxially near blade junction; blade linear, lanceolate, elliptic, or wider leaves pandurate and unequally 4-lobed, occasionally polymorphic on single plants, 15–250 × 4–40 mm, base acute to cuneate, margins subulately glandular-serrulate distally, or sparsely glandular and subentire, hirtellous to glabrate, flat to revolute, apex acute to cuneate, abaxial surface sparsely pilose or glabrate, adaxial surface glabrous or sparsely puberulent; venation pinnate, midvein prominent. |
petiole 0–1 mm; blade oblanceolate to obovate, 8–20 mm × 3–7 mm, base truncate or cuneate, margins entire, apex obtuse to rounded, minutely mucronate, surfaces puberulent or glabrous; venation inconspicuous, only midvein prominent. |
Involucre | campanulate, occasionally broadly so, 1.8–2.8 × 2.2–2.8 mm, glabrous; involucral lobes triangularly 3–5 lobed; gland 1, yellow-green, sessile to substipitate and narrowly to broadly attached, 1–1.4 × 0.9–1.6 mm, opening oblong (flattened without pressing), without annular rim, glabrous; appendages absent. |
cupulate, 2–2.2 × 1.3–1.8 mm, glabrous; glands 4, oblong to broadly ovate, usually truncate, 0.5–0.8 × 1–1.6 mm, margins irregularly crenate to strongly dentate; horns absent or usually divergent or straight, 0.1–0.3 mm, usually slightly longer than, occasionally equaling, teeth on gland margin. |
Staminate flowers | 7–20. |
10–20. |
Pistillate flowers | ovary glabrous; styles 1.6 mm, 2-fid nearly entire length. |
ovary glabrous or puberulent; styles 0.7–1 mm, 2-fid. |
Capsules | green, depressed-globose to ellipsoid, 2.8–3.2 × 4–4.5 mm, 3-lobed, glabrous; columella 2–2.7 mm. |
ovoid, 3.5–4 × 4–4.5 mm, 3-lobed; cocci rounded, smooth to slightly rugose, glabrous; columella 3.2–3.5 mm. |
Seeds | black to ashy gray or light brown, cylindric to ovoid, rounded in cross section, 2.3–3.1 × 1.9–2.5 mm, uniformly tuberculate or tubercles arranged in median, transverse ridge in cylindric seeds; caruncle absent. |
gray to dark gray, truncate-oblong to truncate-ovoid, 2.8–3 × 1.7–2 mm, irregularly pitted; caruncle conic, 0.6 × 0.7 mm. |
Cyathial | arrangement: terminal pleiochasial branches (1–)3, 1–2-branched; pleiochasial bracts 2–3(–4), often as tight, involucrate whorl, usually green with white, pink, or red at base, occasionally distal bracts wholly white, pink, or red, rarely all bracts wholly green, similar in shape and size to distal leaves; dichasial bracts often colored, similar in shape and size to distal stem leaves or highly reduced. |
arrangement: terminal pleiochasial branches 3–5, each 1–4 times 2-branched; pleiochasial bracts ovate to broadly ovate or oblanceolate, wider than distal leaves; dichasial bracts distinct, rounded, oblanceolate, or subreniform, base cuneate or obtuse, margins entire, apex obtuse, slightly mucronate; axillary cymose branches 0–4. |
Cyathia | peduncle 1.6–2.8 mm. |
peduncle 0.3–0.9 mm. |
2n | = 28, 56. |
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Euphorbia cyathophora |
Euphorbia lurida |
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Phenology | Flowering and fruiting late spring–fall. | Flowering and fruiting spring–summer. |
Habitat | Bottomland forests, stream and river banks, bases of bluffs, fallow fields, roadsides, open disturbed areas. | Open pine-oak forests, dry slopes and canyons. |
Elevation | 0–1800 m. (0–5900 ft.) | 1300–2800 m. (4300–9200 ft.) |
Distribution |
AL; AR; CA; FL; GA; IA; IL; IN; KS; KY; LA; MD; MN; MO; MS; NC; NE; NM; OH; OK; SC; SD; TN; TX; UT; VA; WI; Mexico; Central America; South America; West Indies [Introduced in Eurasia, Africa, Indian Ocean Islands, Pacific Islands, Australia]
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AZ; CA; NM; NV; UT; Mexico (Baja California)
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Discussion | Euphorbia cyathophora is native to the midwestern and southeastern United States, Mexico, the West Indies, and northern South America. Leaf shape can be polymorphic on individuals of this species, but not to the extent as in E. heterophylla. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia lurida has been treated as a complex of several taxa in the past, but only a single, broadly-defined species is recognized here. This species is variable in both the pubescence and shape of the bracts subtending the cyathia and also in the degree of crenation of the gland margin. In the northern part of its range, E. lurida appears to intergrade with E. brachycera, and it can be difficult to distinguish these two species in northern Arizona and New Mexico. A report of the species from Sonora, Mexico, based on a single immature collection (V. W. Steinmann and R. S. Felger 1997) has not been verified. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 319. | FNA vol. 12, p. 305. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Poinsettia cyathophora | E. palmeri, E. palmeri var. subpubens, E. subpubens, Tithymalus luridus, T. palmeri, T. subpubens |
Name authority | Murray: Commentat. Soc. Regiae Sci. Gott. 7: 81, plate 1. (1786) | Engelmann: in J. C. Ives, Rep. Colorado R. 4: 26. (1861) |
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