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fire on the mountain, painted leaf, painted poinsettia

bristlecup sandmat

Habit Herbs, annual, with spreading taproots. Herbs, perennial, with woody, thickened taproot.
Stems

erect or ascending, 20–100 cm, glabrous, sparsely pilose, or puberulent;

branches ± straight.

usually erect, rarely slightly decumbent, often densely clustered from top of woody crown, 3–15 cm, glabrous.

Leaves

usually alternate, occasionally opposite proximally;

petiole 2–20 mm, glabrous or pilose, or often hispid abaxially near blade junction;

blade linear, lanceolate, elliptic, or wider leaves pandurate and unequally 4-lobed, occasionally polymorphic on single plants, 15–250 × 4–40 mm, base acute to cuneate, margins subulately glandular-serrulate distally, or sparsely glandular and subentire, hirtellous to glabrate, flat to revolute, apex acute to cuneate, abaxial surface sparsely pilose or glabrate, adaxial surface glabrous or sparsely puberulent;

venation pinnate, midvein prominent.

opposite;

stipules distinct, narrowly linear, usually entire, 0.5–1 mm, glabrous;

petiole 0.5–1 mm, glabrous;

blade ovate to lanceolate or oblong- or linear-lanceolate, 3–11 × 0.8–3(–5) mm, base slightly asymmetric, short-tapered, occasionally one side slightly rounded, margins entire, apex acute or short-acuminate, surfaces glabrous;

only midvein conspicuous.

Involucre

campanulate, occasionally broadly so, 1.8–2.8 × 2.2–2.8 mm, glabrous;

involucral lobes triangularly 3–5 lobed;

gland 1, yellow-green, sessile to substipitate and narrowly to broadly attached, 1–1.4 × 0.9–1.6 mm, opening oblong (flattened without pressing), without annular rim, glabrous;

appendages absent.

campanulate to turbinate, 0.8–1.4 × 0.8–1 mm, glabrous;

glands 4, yellow-brown to reddish, concave or convex, elliptic or oval, 0.2–0.4 × 0.4–0.6 mm;

appendages absent or white, lanceolate-deltate to straplike, 0.2–1.1 × 0.2–0.9 mm, distal margin entire, crenate, or deeply cleft or divided.

Staminate flowers

7–20.

25–35.

Pistillate flowers

ovary glabrous;

styles 1.6 mm, 2-fid nearly entire length.

ovary glabrous;

styles 0.3–0.4 mm, 2-fid 1/2 length.

Capsules

green, depressed-globose to ellipsoid, 2.8–3.2 × 4–4.5 mm, 3-lobed, glabrous;

columella 2–2.7 mm.

depressed-ovoid to depressed-globose, 1.7–2.1 × 1.6–2.4 mm, glabrous;

columella 1.2–1.8 mm.

Seeds

black to ashy gray or light brown, cylindric to ovoid, rounded in cross section, 2.3–3.1 × 1.9–2.5 mm, uniformly tuberculate or tubercles arranged in median, transverse ridge in cylindric seeds;

caruncle absent.

white, ovoid-pyramidal, prominently 4-angled in cross section, 1.6–2 × 1–1.2 mm, smooth to slightly wrinkled.

Cyathial

arrangement: terminal pleiochasial branches (1–)3, 1–2-branched;

pleiochasial bracts 2–3(–4), often as tight, involucrate whorl, usually green with white, pink, or red at base, occasionally distal bracts wholly white, pink, or red, rarely all bracts wholly green, similar in shape and size to distal leaves;

dichasial bracts often colored, similar in shape and size to distal stem leaves or highly reduced.

Cyathia

peduncle 1.6–2.8 mm.

solitary at distal nodes;

peduncle 0.8–1.3 mm.

2n

= 28, 56.

Euphorbia cyathophora

Euphorbia chaetocalyx

Phenology Flowering and fruiting late spring–fall.
Habitat Bottomland forests, stream and river banks, bases of bluffs, fallow fields, roadsides, open disturbed areas.
Elevation 0–1800 m. (0–5900 ft.)
Distribution
from FNA
AL; AR; CA; FL; GA; IA; IL; IN; KS; KY; LA; MD; MN; MO; MS; NC; NE; NM; OH; OK; SC; SD; TN; TX; UT; VA; WI; Mexico; Central America; South America; West Indies [Introduced in Eurasia, Africa, Indian Ocean Islands, Pacific Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; NM; TX; n Mexico
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia cyathophora is native to the midwestern and southeastern United States, Mexico, the West Indies, and northern South America. Leaf shape can be polymorphic on individuals of this species, but not to the extent as in E. heterophylla.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Varieties 2 (2 in the flora).

Euphorbia chaetocalyx is similar to E. fendleri but can generally be distinguished from that species by its narrow, acute leaves and ± erect stems. Some authors have used the presence or absence and shape of the involucral gland appendages to help separate E. chaetocalyx from E. fendleri, but those characters appear highly variable and of little taxonomic utility. Some individuals from western Texas (Culberson and El Paso counties) and southern New Mexico appear intermediate with E. fendleri. The specific epithet of E. chaetocalyx refers to the bristly perianthlike segments that subtend the ovary, but these structures are found intermittently in both E. chaetocalyx and E. fendleri.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Key
1. Leaf blades ovate to lanceolate or oblong- or linear-lanceolate; involucral gland appendages absent or lanceolate-deltate, entire, crenate, or deeply cleft; widely distributed in Arizona, New Mexico, w Texas.
var. chaetocalyx
1. Leaf blades narrowly-ovate to linear-lanceolate; involucral gland appendages straplike, divided into 3–5 linear segments; cliffs and rocks of Boquillas Canyon, sw Texas.
var. triligulata
Source FNA vol. 12, p. 319. FNA vol. 12, p. 263.
Parent taxa Euphorbiaceae > Euphorbia > sect. Poinsettia Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Subordinate taxa
E. chaetocalyx var. chaetocalyx, E. chaetocalyx var. triligulata
Synonyms Poinsettia cyathophora E. fendleri var. chaetocalyx, Chamaesyce chaetocalyx
Name authority Murray: Commentat. Soc. Regiae Sci. Gott. 7: 81, plate 1. (1786) (Boissier) Tidestrom: Proc. Biol. Soc. Wash. 48: 40. (1935)
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