Euphorbia cuphosperma |
Euphorbia serpens |
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hairy-fruit spurge |
creeping spurge, Matted sandmat, Matted spurge, round-leafed spurge |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with slender taproot. |
Stems | erect, 13–20 cm, both pilose to villous and densely strigillose; branches ± straight. |
prostrate, frequently mat-forming and rooting at nodes, 15–50 cm, glabrous. |
Leaves | usually opposite, occasionally alternate at distal nodes; petiole 3–15 mm, pilose; blade narrowly to broadly elliptic, or lanceolate to ovate, 30–80 × 10–15 mm, base cuneate to rounded, margins coarsely crenate-dentate, strigose, revolute to nearly flat, apex broadly acute to acuminate, or obtuse, abaxial surface pilose, adaxial surface sparsely strigose-hirsute; venation pinnate, midvein prominent. |
opposite; stipules connate into conspicuous, deltate scale, white to pink, membranaceous, 0.5–1(–1.2) mm, glabrous; petiole less than 1 mm, glabrous; blade ovate, oblong or orbiculate, 2–7(–9) × 2–6 mm, base asymmetric, rounded to subcordate, margins entire, apex rounded, surfaces without red blotch, glabrous; usually only midvein conspicuous. |
Involucre | campanulate to slightly cylindric, 2.3 × 1.2 mm, glabrous; involucral lobes divided into several linear, smooth lobes; gland 1, yellow-green, stipitate, clavate, 1–1.2 × 0.8–0.9 mm, opening bilabiate and oblong, glabrous; appendages absent. |
campanulate to turbinate, 0.3–0.7(–1) × 0.4–0.6 mm, glabrous; glands 4, yellow, oblong, 0.1 × 0.2 mm; appendages white to pinkish, equal, forming narrow rim at edge of gland, 0.1–0.2 × 0.2–0.3 mm, distal margin entire or crenulate. |
Staminate flowers | 3–5. |
5–10. |
Pistillate flowers | ovary pilose; styles 1.2 mm, 2-fid nearly entire length. |
ovary glabrous; styles 0.2 mm, 2-fid 1/2 to nearly entire length. |
Capsules | broadly ovoid, 2.2–3 × 1.9–2.7 mm, 3-lobed, pilose; columella 2–2.4 mm. |
broadly ovoid, oblate, or subglobose, 1.3–1.4 × 1.3–1.7 mm, glabrous; columella (0.9–)1–1.2 mm. |
Seeds | gray-brown to pale gray, pyramidally ovoid, angular in cross section, 2.3–2.6 × 2.4–2.6 mm, coarsely tuberculate, tubercles in 2 transverse rows; caruncle 0.2–0.4 mm. |
white to gray or light pink, ovoid, bluntly 3–4-angled in cross section, 0.7–1.1 × 0.4–0.7 mm, smooth. |
Cyathial | arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts highly reduced. |
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Cyathia | peduncle 0.5–0.8 mm. |
solitary at distal nodes; peduncle 0.5–1(–2.5) mm. |
2n | = 56. |
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Euphorbia cuphosperma |
Euphorbia serpens |
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Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting year-round (in warmer areas) or summer (in temperate regions). |
Habitat | Open montane and canyon forests, pinyon-juniper forests, montane grasslands, stream beds, disturbed habitats. | Mostly sandy or well-drained soils, desert scrub, coastal scrub, chaparral, oak and juniper woodlands, sand dunes, riparian forests, mesquite grasslands, prairies, coniferous and deciduous hardwood forests, disturbed areas. |
Elevation | 800–2000 m. (2600–6600 ft.) | 0–2000 m. (0–6600 ft.) |
Distribution |
AZ; NM; Mexico; Central America (Guatemala)
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AL; AR; AZ; CA; CO; FL; GA; IA; IL; IN; KS; KY; LA; MI; MO; MS; MT; ND; NE; NJ; NM; OH; OK; PA; SD; TN; TX; UT; VT; WY; ON; SK; Mexico; Central America; South America; West Indies; Bermuda [Introduced in Europe, Asia, Africa, Australia]
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Discussion | Euphorbia serpens is one of the most widespread species of the genus in the New World. While it may be indigenous to a portion of the flora area, probably in the warmer, southern part of its range, it is weedy and has likely been introduced in many parts of the flora area, such as Canada and the eastern United States. It is also widely distributed in the Old World, where it is certainly introduced. The strictly prostrate habit with stems rooting at the nodes is characteristic. Euphorbia serpens is often confused with E. albomarginata, a species distributed in the southwestern United States; in addition to the features mentioned in the key, an easy and reliable way to distinguish between them is by the size of the involucral gland appendages: those of E. albomarginata are conspicuous to the naked eye, whereas those of E. serpens are inconspicuous. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 12, p. 319. | FNA vol. 12, p. 286. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. dentata var. cuphosperma, Poinsettia cuphosperma | Chamaesyce serpens |
Name authority | (Engelmann) Boissier: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 73. (1862) | Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 2(fol.): 41; 2(qto.): 52. (1817) |
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