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hairy-fruit spurge
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revolute spurge, threadstem sandmat, threadstem spurge
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| Herbs, annual, with taproot. |
Herbs, annual, with slender taproot. |
erect, 13–20 cm, both pilose to villous and densely strigillose; branches ± straight. |
erect, 5–25 cm, glabrous. |
usually opposite, occasionally alternate at distal nodes; petiole 3–15 mm, pilose; blade narrowly to broadly elliptic, or lanceolate to ovate, 30–80 × 10–15 mm, base cuneate to rounded, margins coarsely crenate-dentate, strigose, revolute to nearly flat, apex broadly acute to acuminate, or obtuse, abaxial surface pilose, adaxial surface sparsely strigose-hirsute; venation pinnate, midvein prominent. |
opposite; stipules distinct, subulate-filiform, entire, 0.5–0.9 mm, glabrous; petiole 0.6–1.1 mm, glabrous; blade linear, 6–27 × 0.6–1.2 mm, base symmetric, attenuate, margins entire, revolute, apex acute, sometimes mucronate, surfaces glabrous; only midvein conspicuous. |
campanulate to slightly cylindric, 2.3 × 1.2 mm, glabrous; involucral lobes divided into several linear, smooth lobes; gland 1, yellow-green, stipitate, clavate, 1–1.2 × 0.8–0.9 mm, opening bilabiate and oblong, glabrous; appendages absent. |
obconic to campanulate, 0.7–0.9 × 0.5–0.7 mm, glabrous; glands 4, pink to dark purple, nearly circular to oblong or reniform, 0.1 × 0.1–0.2 mm; appendages white, oblong, nearly circular, ovate, deltate, or forming thin margin around gland, rarely absent, (0–)0.1–0.2 × (0–)0.1–0.3 mm, distal margin entire. |
3–5. |
5–10. |
ovary pilose; styles 1.2 mm, 2-fid nearly entire length. |
ovary glabrous; styles 0.2–0.3 mm, unbranched. |
broadly ovoid, 2.2–3 × 1.9–2.7 mm, 3-lobed, pilose; columella 2–2.4 mm. |
ovoid to subglobose, 1.5–1.8 × 1.6–1.8 mm, glabrous; columella 1.1–1.5 mm. |
gray-brown to pale gray, pyramidally ovoid, angular in cross section, 2.3–2.6 × 2.4–2.6 mm, coarsely tuberculate, tubercles in 2 transverse rows; caruncle 0.2–0.4 mm. |
whitish, brick red, light gray, or light brown, narrowly to broadly ovoid, 4-angled in cross section, 0.9–1.4 × 0.7–1 mm, nearly smooth, rugulose, with faint transverse ridges, or with 2–3 well-defined transverse ridges separated by shallow depressions. |
arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts highly reduced. |
|
peduncle 0.5–0.8 mm. |
solitary at distal nodes; peduncles absent or to 1.5 mm. |
= 56. |
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| Flowering and fruiting summer–fall. |
Flowering and fruiting summer–late fall. |
| Open montane and canyon forests, pinyon-juniper forests, montane grasslands, stream beds, disturbed habitats. |
Desert scrub, sagebrush scrub, juniper woodlands, Joshua tree-pinyon pine woodlands, oak woodlands, grasslands, chaparral, pine-oak forests. |
| 800–2000 m. (2600–6600 ft.) |
600–2500 m. (2000–8200 ft.) |
|
AZ; NM; Mexico; Central America (Guatemala)
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AZ; CA; CO; NM; NV; TX; UT; Mexico (Baja California, Chihuahua, Coahuila, Sonora, Zacatecas)
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Euphorbia revoluta is a distinctive, easily recognizable species by virtue of the combination of unbranched styles and linear leaves with symmetric bases. The species ranges through northern Mexico into the southwestern United States and is composed of three well-marked, geographically distinct races that can be distinguished by their seeds. The first of these races occurs primarily in pine-oak forest of the Sierra Madre Occidental of Mexico (Chihuahua and Sonora) and barely enters the flora area in the Huachuca Mountains of southeastern Arizona; the seeds are brick red and nearly smooth. The second race is widespread in the Chihuahuan Desert from northern Mexico to southeastern New Mexico and southwestern Texas; its seeds are whitish to light gray and possess two or three well-defined transverse ridges separated by shallow depressions. The third race corresponds to the type collection and is characterized by grayish white to light brown seeds that are rugulose or with faint transverse ridges; it occurs in northwestern Mexico (Baja California and Sonora) and throughout the arid southwestern United States. It is probable that further study will justify the taxonomic segregation of these races as either distinct species or subspecies. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 12, p. 319. |
FNA vol. 12, p. 286. |
| Euphorbiaceae > Euphorbia > sect. Poinsettia |
Euphorbiaceae > Euphorbia > sect. Anisophyllum |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
| E. dentata var. cuphosperma, Poinsettia cuphosperma |
Chamaesyce revoluta |
| (Engelmann) Boissier: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 73. (1862) |
Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 186. (1859) |
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