Euphorbia cuphosperma |
Euphorbia polycarpa |
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hairy-fruit spurge |
smallseed sandmat |
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Habit | Herbs, annual, with taproot. | |
Stems | erect, 13–20 cm, both pilose to villous and densely strigillose; branches ± straight. |
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Leaves | usually opposite, occasionally alternate at distal nodes; petiole 3–15 mm, pilose; blade narrowly to broadly elliptic, or lanceolate to ovate, 30–80 × 10–15 mm, base cuneate to rounded, margins coarsely crenate-dentate, strigose, revolute to nearly flat, apex broadly acute to acuminate, or obtuse, abaxial surface pilose, adaxial surface sparsely strigose-hirsute; venation pinnate, midvein prominent. |
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Involucre | campanulate to slightly cylindric, 2.3 × 1.2 mm, glabrous; involucral lobes divided into several linear, smooth lobes; gland 1, yellow-green, stipitate, clavate, 1–1.2 × 0.8–0.9 mm, opening bilabiate and oblong, glabrous; appendages absent. |
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Staminate flowers | 3–5. |
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Pistillate flowers | ovary pilose; styles 1.2 mm, 2-fid nearly entire length. |
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Capsules | broadly ovoid, 2.2–3 × 1.9–2.7 mm, 3-lobed, pilose; columella 2–2.4 mm. |
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Seeds | gray-brown to pale gray, pyramidally ovoid, angular in cross section, 2.3–2.6 × 2.4–2.6 mm, coarsely tuberculate, tubercles in 2 transverse rows; caruncle 0.2–0.4 mm. |
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Cyathial | arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts highly reduced. |
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Cyathia | peduncle 0.5–0.8 mm. |
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Euphorbia | polycarpa is a highly variable species distributed throughout the Baja California peninsula, eastern Sonora, and the arid southwestern United States from southwestern Arizona to southern Nevada and central California. |
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l | . |
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c | . |
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Wheeler | (1941) divided the species into seven varieties, the majority of which occur in Baja California Sur. |
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He | reported two of these for the United States: var. hirtella and var. polycarpa. |
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Variety | hirtella, as the name suggests, was applied to hairy plants. |
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However | , it is here treated as a synonym of var. polycarpa, because variation in pubescence shows no geographic segregation and ranges along a continuum from glabrous or sparsely to densely hairy, and because glabrous and hairy branches can occur on the same individual. |
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2n | = 56. |
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Euphorbia cuphosperma |
Euphorbia polycarpa |
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Phenology | Flowering and fruiting summer–fall. | |
Habitat | Open montane and canyon forests, pinyon-juniper forests, montane grasslands, stream beds, disturbed habitats. | |
Elevation | 800–2000 m. (2600–6600 ft.) | |
Distribution |
AZ; NM; Mexico; Central America (Guatemala)
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AZ; CA; NV; nw Mexico
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Discussion | Varieties 5 (1 in the flora). Some of the varieties occurring in Mexico are highly divergent and appear to be sufficiently distinct to merit recognition as species. However, within the flora area, Euphorbia polycarpa is relatively uniform, and the only noteworthy variation involves the size of the involucral gland appendages. In the portion of the California Floristic Province occupied by E. polycarpa (Los Angeles, Orange, western Riverside, western San Diego, and Ventura counties) the appendages are conspicuously larger than those of plants throughout the remainder of its range in the southwestern United States (0.4–1.2 versus 0.1–0.3 mm). Large-appendaged plants are also common in Baja California and correspond well with the type collection. The taxonomic significance of this variation is not clear at this time, and the small-appendaged plants may merit segregation as an infraspecific taxon. However, no formal changes are proposed, awaiting a comprehensive review of the species throughout its range. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 12, p. 319. | FNA vol. 12, p. 283. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Subordinate taxa | ||
Synonyms | E. dentata var. cuphosperma, Poinsettia cuphosperma | Chamaesyce polycarpa |
Name authority | (Engelmann) Boissier: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 73. (1862) | Bentham: Bot. Voy. Sulphur, 50. (1844) |
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