Euphorbia cuphosperma |
Euphorbia ouachitana |
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hairy-fruit spurge |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with taproot. |
Stems | erect, 13–20 cm, both pilose to villous and densely strigillose; branches ± straight. |
erect-ascending, often basally decumbent, often branched near base, 12–28 cm, glabrous. |
Leaves | usually opposite, occasionally alternate at distal nodes; petiole 3–15 mm, pilose; blade narrowly to broadly elliptic, or lanceolate to ovate, 30–80 × 10–15 mm, base cuneate to rounded, margins coarsely crenate-dentate, strigose, revolute to nearly flat, apex broadly acute to acuminate, or obtuse, abaxial surface pilose, adaxial surface sparsely strigose-hirsute; venation pinnate, midvein prominent. |
petiole 0–3 mm; blade broadly oblanceolate to subspatulate, or proximalmost often orbiculate, 3–20 × 3–9 mm, base attenuate, margins entire, apex rounded to emarginate, surfaces glabrous; venation pinnate, midvein prominent. |
Involucre | campanulate to slightly cylindric, 2.3 × 1.2 mm, glabrous; involucral lobes divided into several linear, smooth lobes; gland 1, yellow-green, stipitate, clavate, 1–1.2 × 0.8–0.9 mm, opening bilabiate and oblong, glabrous; appendages absent. |
infundibular, 1.3–1.6 × 1–1.2 mm, glabrous; glands 4, crescent-shaped; 0.4–0.5 × 0.8–1 mm; horns divergent, 0.2–0.4 mm. |
Staminate flowers | 3–5. |
15–20. |
Pistillate flowers | ovary pilose; styles 1.2 mm, 2-fid nearly entire length. |
ovary glabrous; styles 0.8 mm, 2-fid. |
Capsules | broadly ovoid, 2.2–3 × 1.9–2.7 mm, 3-lobed, pilose; columella 2–2.4 mm. |
ovoid-globose, 2.6–2.7 × 2.5–2.7 mm, slightly lobed; cocci rounded to ± flattened, smooth, glabrous; columella 2–2.1 mm. |
Seeds | gray-brown to pale gray, pyramidally ovoid, angular in cross section, 2.3–2.6 × 2.4–2.6 mm, coarsely tuberculate, tubercles in 2 transverse rows; caruncle 0.2–0.4 mm. |
dark brown, oblong-ovoid, 1.5–2 × 1–1.6 mm, with deep, rounded pits in 3–4 regular vertical rows; caruncle reniform-ovate, conic, 0.5–0.7 × 0.5–0.8 mm. |
Cyathial | arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts highly reduced. |
arrangement: terminal pleiochasial branches 3, 1–3 times 2-branched; pleiochasial bracts ovate-deltate to subrhombic-ovate, shorter and wider than distal leaves; dichasial bracts connate 1/4 length, not imbricate, broadly deltate to subreniform, base truncate to broadly obtuse, margins entire, apex obtuse to bluntly acuminate; axillary cymose branches 1–5. |
Cyathia | peduncle 0.5–0.8 mm. |
peduncle 0.4–0.7 mm. |
2n | = 56. |
= 26. |
Euphorbia cuphosperma |
Euphorbia ouachitana |
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Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting spring. |
Habitat | Open montane and canyon forests, pinyon-juniper forests, montane grasslands, stream beds, disturbed habitats. | Semiopen forests, bluffs and ledges, stream banks, glades. |
Elevation | 800–2000 m. (2600–6600 ft.) | 100–400 m. (300–1300 ft.) |
Distribution |
AZ; NM; Mexico; Central America (Guatemala)
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AR; MO; OK; TN; TX |
Discussion | Euphorbia ouachitana is restricted to semiopen forests and woodlands in the south-central United States. It is similar to E. commutata and has been most commonly identified as that species in the past. However, it differs in its consistently brown seeds that have pits in regular, vertical rows. The two species also differ in E. commutata’s numerous, proximal, long-petiolate elliptic leaves. The proximal leaves of E. ouachitana are spatulate with an orbiculate blade and petiolelike base. Aside from the restricted distribution of E. ouachitana in Missouri and Tennessee, the ranges of E. ouachitana and E. commutata do not overlap (see M. H. Mayfield 2013 for a detailed discussion of the distribution). Euphorbia ouachitana is most common in the Ouachita Mountains from southeastern Oklahoma to Hot Springs County, Arkansas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 12, p. 319. | FNA vol. 12, p. 306. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | E. dentata var. cuphosperma, Poinsettia cuphosperma | |
Name authority | (Engelmann) Boissier: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 73. (1862) | Mayfield: J. Bot. Res. Inst. Texas 7: 642, figs. 2[row 2, right], 6. (2013) |
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