Euphorbia cuphosperma |
Euphorbia nutans |
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hairy-fruit spurge |
eyebane, eyebane sandmat, eyebane spurge, nodding or upright spotted spurge, nodding spurge |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with taproot. |
Stems | erect, 13–20 cm, both pilose to villous and densely strigillose; branches ± straight. |
usually ascending, occasionally erect, often arched at tips, 20–80 cm, sparsely to moderately pilose to villous or with short, incurved hairs, hairs often concentrated at nodes and distally, occasionally in 2 bands along opposite sides of stem. |
Leaves | usually opposite, occasionally alternate at distal nodes; petiole 3–15 mm, pilose; blade narrowly to broadly elliptic, or lanceolate to ovate, 30–80 × 10–15 mm, base cuneate to rounded, margins coarsely crenate-dentate, strigose, revolute to nearly flat, apex broadly acute to acuminate, or obtuse, abaxial surface pilose, adaxial surface sparsely strigose-hirsute; venation pinnate, midvein prominent. |
opposite; stipules usually distinct, sometimes connate basally on one side of stem, small scales, irregularly toothed, fringed, or divided, 1–1.5 mm, sparsely to moderately villous distally; petiole 0.3–1.6 mm, moderately pilose to villous; blade oblong to oblong-lanceolate, 8–40 × 3–12 mm, base asymmetric, one side usually angled or rounded, other side ± truncate to cordate-auriculate, margins serrulate, apex angled with blunt tip, abaxial surface pale green or faintly to strongly reddish tinged, adaxial surface usually reddish-mottled or with conspicuous reddish spot, both surfaces usually sparsely to moderately pilose, especially toward base, sometimes glabrous; 3–5-veined from base, pinnate distally, veins faint. |
Involucre | campanulate to slightly cylindric, 2.3 × 1.2 mm, glabrous; involucral lobes divided into several linear, smooth lobes; gland 1, yellow-green, stipitate, clavate, 1–1.2 × 0.8–0.9 mm, opening bilabiate and oblong, glabrous; appendages absent. |
narrowly obconic, 0.5–1 × 0.3–0.7 mm, glabrous; glands 4, usually green, sometimes reddish purple, oblong to nearly circular, 0.2–0.4 × 0.3–0.5 mm; appendages white or pinkish, ovate to broadly elliptic, 0.2–1 × 0.2–1.5 mm, distal margin entire. |
Staminate flowers | 3–5. |
5–28. |
Pistillate flowers | ovary pilose; styles 1.2 mm, 2-fid nearly entire length. |
ovary glabrous; styles 0.6–2.5 mm, 2-fid 1/2 length. |
Capsules | broadly ovoid, 2.2–3 × 1.9–2.7 mm, 3-lobed, pilose; columella 2–2.4 mm. |
ovoid, 1.6–2.3 × 1.5–2.4 mm, glabrous; columella 1.4–1.6 mm. |
Seeds | gray-brown to pale gray, pyramidally ovoid, angular in cross section, 2.3–2.6 × 2.4–2.6 mm, coarsely tuberculate, tubercles in 2 transverse rows; caruncle 0.2–0.4 mm. |
dark brown, sometimes with thin, white coating (often more persistent along angles than faces), elliptic-ovoid to ovoid, rounded-angular in cross section, 1–1.6 × 0.5–0.8 mm, surface finely and irregularly wrinkled, sometimes faintly so, or with indistinct, shallow, rounded cross ridges. |
Cyathial | arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts highly reduced. |
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Cyathia | peduncle 0.5–0.8 mm. |
solitary at distal nodes or in small, cymose clusters at branch tips; peduncle 0.5–2.5 mm. |
2n | = 56. |
= 12, 14, 22. |
Euphorbia cuphosperma |
Euphorbia nutans |
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Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting spring–early fall. |
Habitat | Open montane and canyon forests, pinyon-juniper forests, montane grasslands, stream beds, disturbed habitats. | Stream banks, pond edges, disturbed portions of upland prairies, mesic to dry upland forest openings, pastures, fallow fields, railroads, roadsides, gardens, disturbed areas. |
Elevation | 800–2000 m. (2600–6600 ft.) | 0–1600 m. (0–5200 ft.) |
Distribution |
AZ; NM; Mexico; Central America (Guatemala)
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AL; AR; CA; CT; DE; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MN; MO; MS; NC; ND; NE; NH; NJ; NM; NY; OH; OK; PA; RI; SC; SD; TN; TX; VA; VT; WI; WV; WY; ON; QC; Mexico; Central America; South America; West Indies [Introduced in Eurasia]
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Discussion | Euphorbia nutans is probably native to at least central and eastern North America, but given its strongly weedy tendencies, it is difficult to know where it may be adventive in parts of the flora area. It is certainly introduced where it occurs in the Old World and probably in South America as well. D. G. Burch (1966) discussed the problems of assigning names to the four main entities in this nomenclatural complex (E. hypericifolia, E. hyssopifolia, E. lasiocarpa, and E. nutans) and determined that the oldest valid name for the relatively robust, temperate North American plants with ascending stems is E. nutans. See the treatment of 56. E. maculata for a discussion of the misapplication of that name to E. nutans. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 12, p. 319. | FNA vol. 12, p. 278. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Poinsettia | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | E. dentata var. cuphosperma, Poinsettia cuphosperma | Chamaesyce lansingii, C. nutans, E. lansingii, E. preslii |
Name authority | (Engelmann) Boissier: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 73. (1862) | Lagasca: Gen. Sp. Pl., 17. (1816) |
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