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hairy-fruit spurge

caper spurge, euphorbe épurge, gopher plant, gopher spurge, mole plant

Habit Herbs, annual, with taproot. Herbs, annual or biennial, with taproot.
Stems

erect, 13–20 cm, both pilose to villous and densely strigillose;

branches ± straight.

erect, unbranched or branched, 200 cm, glabrous, glaucous.

Leaves

usually opposite, occasionally alternate at distal nodes;

petiole 3–15 mm, pilose;

blade narrowly to broadly elliptic, or lanceolate to ovate, 30–80 × 10–15 mm, base cuneate to rounded, margins coarsely crenate-dentate, strigose, revolute to nearly flat, apex broadly acute to acuminate, or obtuse, abaxial surface pilose, adaxial surface sparsely strigose-hirsute;

venation pinnate, midvein prominent.

opposite, decussate;

petiole absent;

blade linear to oblong-lanceolate, 30–120 × 3–25 mm, base acute, rounded, cordate or clasping, margins entire, apex acute or subobtuse, sometimes mucronate, surfaces glabrous, abaxial ± glaucous;

venation pinnate, midvein prominent.

Involucre

campanulate to slightly cylindric, 2.3 × 1.2 mm, glabrous;

involucral lobes divided into several linear, smooth lobes;

gland 1, yellow-green, stipitate, clavate, 1–1.2 × 0.8–0.9 mm, opening bilabiate and oblong, glabrous;

appendages absent.

campanulate, 1.2–2.3 × 1.4–2.5 mm, glabrous;

glands 4, elliptic, 0.3–0.6 × 1–1.3 mm;

horns divergent, thick, tips rounded, dilated, 0.5–1.4 mm.

Staminate flowers

3–5.

25–30.

Pistillate flowers

ovary pilose;

styles 1.2 mm, 2-fid nearly entire length.

ovary glabrous;

styles 0.7–2.2 mm, 2-fid.

Capsules

broadly ovoid, 2.2–3 × 1.9–2.7 mm, 3-lobed, pilose;

columella 2–2.4 mm.

depressed-globose, 9–12 × 12–16 mm, deeply 3-lobed, tardily dehiscent and appearing indehiscent, mesocarp spongy;

cocci rounded, smooth, glabrous;

columella 4–5.2 mm.

Seeds

gray-brown to pale gray, pyramidally ovoid, angular in cross section, 2.3–2.6 × 2.4–2.6 mm, coarsely tuberculate, tubercles in 2 transverse rows;

caruncle 0.2–0.4 mm.

brownish or blackish, oblong, 4.5–6 × 3–4.2 mm, rugose, irregularly reticulate;

caruncle substipitate, hat-shaped, 1.6–2 × 1.2–1.5 mm.

Cyathial

arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched;

pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower;

dichasial bracts highly reduced.

arrangement: terminal pleiochasial branches 2–4, each 1–2 times 2-branched;

pleiochasial bracts cordate-lanceolate, shorter and wider than distal leaves;

dichasial bracts distinct, ovate-oblong to lanceolate, base subcordate, margins entire, apex acute;

axillary cymose branches 0–10.

Cyathia

peduncle 0.5–0.8 mm.

peduncle 0–0.5 mm.

2n

= 56.

Euphorbia cuphosperma

Euphorbia lathyris

Phenology Flowering and fruiting summer–fall. Flowering and fruiting winter–fall.
Habitat Open montane and canyon forests, pinyon-juniper forests, montane grasslands, stream beds, disturbed habitats. Roadsides, cultivated fields, stream banks, waste places.
Elevation 800–2000 m. (2600–6600 ft.) 0–1800 m. (0–5900 ft.)
Distribution
from FNA
AZ; NM; Mexico; Central America (Guatemala)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; CT; ID; IL; MA; MD; MT; NC; OH; OR; PA; SC; TN; TX; VA; WA; WV; BC; ON; QC; Europe; Asia; Africa [Introduced in North America; introduced also in Mexico, South America, Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia lathyris is most likely native to the central and eastern Mediterranean region, but it is widely cultivated and often locally escaped in temperate regions worldwide, as in the flora area.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 319. FNA vol. 12, p. 304.
Parent taxa Euphorbiaceae > Euphorbia > sect. Poinsettia Euphorbiaceae > Euphorbia > subg. Esula
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms E. dentata var. cuphosperma, Poinsettia cuphosperma Galarhoeus lathyris, Tithymalus lathyris
Name authority (Engelmann) Boissier: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 73. (1862) Linnaeus: Sp. Pl. 1: 457. (1753) — (as lathyrus)
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