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hairy-fruit spurge

Arizona sandmat, Arizona spurge

Habit Herbs, annual, with taproot. Herbs, annual or short-lived perennial, with slender to slightly thickened taproot.
Stems

erect, 13–20 cm, both pilose to villous and densely strigillose;

branches ± straight.

erect to ascending, 10–30 cm, uniformly pilose with glistening hairs.

Leaves

usually opposite, occasionally alternate at distal nodes;

petiole 3–15 mm, pilose;

blade narrowly to broadly elliptic, or lanceolate to ovate, 30–80 × 10–15 mm, base cuneate to rounded, margins coarsely crenate-dentate, strigose, revolute to nearly flat, apex broadly acute to acuminate, or obtuse, abaxial surface pilose, adaxial surface sparsely strigose-hirsute;

venation pinnate, midvein prominent.

opposite;

stipules distinct, deltate, (0–)0.1 mm, glabrous or with few scattered hairs;

petiole 0.4–1.5 mm, pilose with glistening hairs;

blade usually ovate, rarely elliptic, 3–11 × 2–7 mm, base asymmetric, one side cuneate to rounded, other side rounded to strongly cordate, margins entire, apex obtuse or acute, surfaces pilose with glistening hairs; 3-veined from base but usually only midvein conspicuous.

Involucre

campanulate to slightly cylindric, 2.3 × 1.2 mm, glabrous;

involucral lobes divided into several linear, smooth lobes;

gland 1, yellow-green, stipitate, clavate, 1–1.2 × 0.8–0.9 mm, opening bilabiate and oblong, glabrous;

appendages absent.

urceolate, 1.1–1.3 × 0.5–0.9 mm, glabrous or pilose with glistening hairs;

glands 4, dark maroon, usually oblong to reniform, rarely almost circular, 0.2 × 0.2–0.4 mm;

appendages white to pink, oblong, flabellate, oblong, or elliptic, 0.3–0.6 × 0.6–0.9 mm, distal margin entire or crenulate.

Staminate flowers

3–5.

5–12.

Pistillate flowers

ovary pilose;

styles 1.2 mm, 2-fid nearly entire length.

ovary pilose, styles 0.5–0.6 mm, 2-fid 1/2 length.

Capsules

broadly ovoid, 2.2–3 × 1.9–2.7 mm, 3-lobed, pilose;

columella 2–2.4 mm.

broadly ovoid to subglobose, 1.4–1.8 mm diam., pilose;

columella 1.1–1.4 mm.

Seeds

gray-brown to pale gray, pyramidally ovoid, angular in cross section, 2.3–2.6 × 2.4–2.6 mm, coarsely tuberculate, tubercles in 2 transverse rows;

caruncle 0.2–0.4 mm.

gray to light brown, ovoid, 4-angled in cross section, 0.9–1.1 × 0.5–0.6 mm, rugose with 2–5 irregular transverse ridges that sometimes pass through abaxial keel.

Cyathial

arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched;

pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower;

dichasial bracts highly reduced.

Cyathia

peduncle 0.5–0.8 mm.

solitary at distal nodes;

peduncle 0.7–3.6 mm.

2n

= 56.

Euphorbia cuphosperma

Euphorbia arizonica

Phenology Flowering and fruiting summer–fall. Flowering and fruiting year-round in response to sufficient moisture.
Habitat Open montane and canyon forests, pinyon-juniper forests, montane grasslands, stream beds, disturbed habitats. Washes and rocky slopes, sometimes on limestone, desert scrub communities often with creosote-bush, riparian forests, mesquite woodlands, oak chaparral.
Elevation 800–2000 m. (2600–6600 ft.) 100–1400 m. (300–4600 ft.)
Distribution
from FNA
AZ; NM; Mexico; Central America (Guatemala)
[WildflowerSearch map]
[BONAP county map]
from FNA
AZ; CA; NM; TX; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, Sonora)
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia arizonica is distinctive and easily recognized by its glistening, translucent hairs that appear somewhat glutinous and are most apparent on the stems.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 319. FNA vol. 12, p. 260.
Parent taxa Euphorbiaceae > Euphorbia > sect. Poinsettia Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms E. dentata var. cuphosperma, Poinsettia cuphosperma Chamaesyce arizonica
Name authority (Engelmann) Boissier: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 73. (1862) Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 186. (1859)
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