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hairy-fruit spurge

Blackfoot sandmat, narrow-leaf spurge

Habit Herbs, annual, with taproot. Herbs, perennial, with moderately to strongly thickened rootstock.
Stems

erect, 13–20 cm, both pilose to villous and densely strigillose;

branches ± straight.

erect, 12–43 cm, uniformly strigose.

Leaves

usually opposite, occasionally alternate at distal nodes;

petiole 3–15 mm, pilose;

blade narrowly to broadly elliptic, or lanceolate to ovate, 30–80 × 10–15 mm, base cuneate to rounded, margins coarsely crenate-dentate, strigose, revolute to nearly flat, apex broadly acute to acuminate, or obtuse, abaxial surface pilose, adaxial surface sparsely strigose-hirsute;

venation pinnate, midvein prominent.

opposite;

stipules distinct, linear-subulate or nodiform to papilliform nodiform to papilliform stipules often reddish brown, 0.1–0.7 mm, strigose;

petiole 0.3–1.2 mm, strigose;

blade: proximal ovate to ovate-elliptic, distal linear to elliptic-linear, 7–41 × 2–5 mm, distal leaf blades more than 6 times as long as wide, base asymmetric, cuneate to rounded, margins entire, often involute on drying, apex acute, surfaces usually short strigose, occasionally glabrous adaxially;

venation pinnate, only midvein conspicuous.

Involucre

campanulate to slightly cylindric, 2.3 × 1.2 mm, glabrous;

involucral lobes divided into several linear, smooth lobes;

gland 1, yellow-green, stipitate, clavate, 1–1.2 × 0.8–0.9 mm, opening bilabiate and oblong, glabrous;

appendages absent.

turbinate to campanulate-turbinate, 1–1.5 × 1–1.4 mm, strigose;

glands 4, green to yellow-green, concave, narrowly oblong, 0.2–0.4 × 0.4–0.7 mm;

appendages white, flabellate, 0.5–1.1 × 0.3–0.5 mm, distal margin shallowly and irregularly toothed.

Staminate flowers

3–5.

16–26.

Pistillate flowers

ovary pilose;

styles 1.2 mm, 2-fid nearly entire length.

ovary strigose;

styles 0.3–0.5 mm, 2-fid at apex to almost 1/2 length.

Capsules

broadly ovoid, 2.2–3 × 1.9–2.7 mm, 3-lobed, pilose;

columella 2–2.4 mm.

broadly ovoid, 2–2.6 × 2.5–3 mm, strigose;

columella 1.6–2.2 mm.

Seeds

gray-brown to pale gray, pyramidally ovoid, angular in cross section, 2.3–2.6 × 2.4–2.6 mm, coarsely tuberculate, tubercles in 2 transverse rows;

caruncle 0.2–0.4 mm.

white, ovoid, 4-angled in cross section, 1.7–2.2 × 1.1–1.2 mm, transversely low-ridged or wrinkled.

Cyathial

arrangement: terminal pleiochasial branches usually 3, occasionally reduced to congested cyme, 1–2-branched;

pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower;

dichasial bracts highly reduced.

Cyathia

peduncle 0.5–0.8 mm.

solitary at distal nodes;

peduncle 1.1–2.4 mm.

2n

= 56.

= 28.

Euphorbia cuphosperma

Euphorbia angusta

Phenology Flowering and fruiting summer–fall. Flowering and fruiting early spring–fall.
Habitat Open montane and canyon forests, pinyon-juniper forests, montane grasslands, stream beds, disturbed habitats. Rocky limestone soils.
Elevation 800–2000 m. (2600–6600 ft.) 400–1200 m. (1300–3900 ft.)
Distribution
from FNA
AZ; NM; Mexico; Central America (Guatemala)
[WildflowerSearch map]
[BONAP county map]
from FNA
TX; Mexico (Coahuila)
[BONAP county map]
Discussion

Euphorbia angusta, which in the flora area is known from the trans-Pecos region to the Edwards Plateau, is easily recognized by its erect habit, linear leaves, and relatively showy involucral gland appendages with toothed margins. The species is closely related to E. acuta and the Mexican endemic E. johnstonii Mayfield (M. H. Mayfield 1991); it is not only morphologically distinctive but is also the only species in sect. Anisophyllum with C3 photosynthesis (R. F. Sage et al. 2011; T. L. Sage et al. 2011; G. L. Webster 1975).

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 319. FNA vol. 12, p. 260.
Parent taxa Euphorbiaceae > Euphorbia > sect. Poinsettia Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms E. dentata var. cuphosperma, Poinsettia cuphosperma Chamaesyce angusta
Name authority (Engelmann) Boissier: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 73. (1862) Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859)
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