Euphorbia crenulata |
Euphorbia vallis-mortae |
|
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Chinese caps, western wood spurge |
Death Valley sandmat |
|
Habit | Herbs, usually biennial, occasionally annual, with taproot. | Herbs, perennial, with thickened, woody taproot. |
Stems | erect, sometimes decumbent at base, unbranched or branched, 12–40 cm, glabrous. |
prostrate to ascending, often mat-forming, 10–45 cm, pilose to villous. |
Leaves | petiole 0–2 mm; blade obovate-spatulate to oblanceolate, 8–22 × 3–10 mm, base broadly attenuate, margins entire or slightly crisped, apex obtuse to ± rounded, minutely apiculate, surfaces glabrous; venation pinnate, midvein prominent. |
opposite; stipules distinct or connate, subulate to filiform, 0.4–1.1 mm, densely tomentose; petiole 0.4–1 mm, pilose to villous; blade suborbiculate to oblong-ovate, 3–8 × 2–6 mm, base slightly asymmetric, rounded, margins entire, apex rounded to obtuse, surfaces pilose to villous; 3-veined at base, midvein conspicuous, venation often obscured by pubescence. |
Involucre | campanulate, 1.8–2.1 × 1.6–1.8 mm, glabrous; glands 4, crescent-shaped, 0.6–1.2 × 1.5–2.3 mm; horns slightly divergent to slightly convergent, 0.4–0.6 mm. |
obconic-campanulate, 1.2–2.3 × 1–1.8 mm, densely pilose to villous; glands 4, yellow to red, subcircular to oblong, 0.2–0.5 × 0.4–0.9 mm; appendages white, flabellate to oblong, 0.1–0.7 × 0.5–1.9 mm, distal margin entire or crenulate, adaxial surface ciliate-puberulent. |
Staminate flowers | 11–18. |
15–22(–50). |
Pistillate flowers | ovary glabrous; styles 0.9–1.4 mm, 2-fid. |
ovary densely pilose; styles 0.4–0.8 mm, 2-fid 1/2 length. |
Capsules | subovoid, 2.5–3 × 3.5–4 mm, 3-lobed; cocci rounded, smooth or puncticulate, glabrous; columella 1.9–2.3 mm. |
ovoid, 1.5–2.2 × 1.8–2.2 mm, tomentose; columella 1.2–1.6 mm. |
Seeds | cream and brown mottled, oblong-ovoid to nearly globose, 2–2.5 × 1.4–1.7 mm, usually irregularly vermiculate-ridged and large-pitted, occasionally tuberculate or nearly smooth; caruncle reniform, conic, 0.5–0.6 × 0.5–0.7 mm. |
white, gray, or light brown, ovoid, sharply 4-angled in cross section, abaxial faces slightly convex, adaxial faces concave, 1.2–1.7 × 0.6–0.9 mm, smooth. |
Cyathial | arrangement: terminal pleiochasial branches 3, each 2-branched; pleiochasial bracts obovate to orbiculate-reniform, wider than distal leaves; dichasial bracts usually connate 1/3–1/2 length (often only on one side), rarely only connate basally, triangular ovate to reniform, base truncate to perfoliate, margins erose-denticulate to subentire, apex rounded to obtuse, rarely apiculate; axillary cymose branches 0–5. |
|
Cyathia | peduncle 0–0.5 mm. |
solitary at distal nodes; peduncle 0.5–1.8 mm. |
Euphorbia crenulata |
Euphorbia vallis-mortae |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting late spring–fall. |
Habitat | Conifer, oak, and mixed forests, coastal scrub, grasslands, barrens and outcrops, roadsides. | Roadsides, desert scrub, streamsides, sandy washes. |
Elevation | 30–1800 m. (100–5900 ft.) | 700–2000 m. (2300–6600 ft.) |
Distribution |
CA; CO; NM; OR
|
CA |
Discussion | Euphorbia crenulata is most common in the central valleys of California and southern Oregon; it occurs disjunctly in southwestern Colorado and northwestern New Mexico. Previous reports from Arizona are based on misidentified specimens. Euphorbia crenulata is closely related to E. commutata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
The specific epithet of Euphorbia vallis-mortae is a misnomer because the species does not occur in Death Valley; instead, it is found at the transition of the northern edge of the Mojave Desert and the foothills of the southern Sierra Nevada in Inyo, Kern, and San Bernardino counties. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 301. | FNA vol. 12, p. 291. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Tithymalus crenulatus | Chamaesyce vallis-mortae |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) | (Millspaugh) J. T. Howell: Madroño 2: 19. (1931) |
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