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Chinese caps, western wood spurge

Habit Herbs, usually biennial, occasionally annual, with taproot. Herbs, annual, with slender taproot.
Stems

erect, sometimes decumbent at base, unbranched or branched, 12–40 cm, glabrous.

prostrate, 4–8 cm, uniformly strigillose.

Leaves

petiole 0–2 mm;

blade obovate-spatulate to oblanceolate, 8–22 × 3–10 mm, base broadly attenuate, margins entire or slightly crisped, apex obtuse to ± rounded, minutely apiculate, surfaces glabrous;

venation pinnate, midvein prominent.

opposite;

stipules distinct, filiform, 0.6–0.9 mm, pilose;

petiole 0.5–0.9 mm, strigillose;

blade ovate to elliptic, often slightly falcate, 5–11 × 2–5 mm, base asymmetric, one side cordate, other round to attenuate, margins sharply serrulate, apex acute, surfaces often with red spot toward middle, abaxial surface strigillose, adaxial surface glabrescent;

only midvein conspicuous.

Involucre

campanulate, 1.8–2.1 × 1.6–1.8 mm, glabrous;

glands 4, crescent-shaped, 0.6–1.2 × 1.5–2.3 mm;

horns slightly divergent to slightly convergent, 0.4–0.6 mm.

obconic, 0.9–1.2 × 0.8–1.2 mm, strigillose;

glands 4, green, yellow, or light pink, circular to oblong, 0.2 × 0.2–0.3 mm;

appendages absent or green, yellow, or light pink, forming narrow margin on distal portion of gland, 0–0.1 × 0.2–0.3 mm, distal margin entire.

Staminate flowers

11–18.

5–8.

Pistillate flowers

ovary glabrous;

styles 0.9–1.4 mm, 2-fid.

ovary strigillose-canescent;

styles 0.3–0.4 mm, unbranched.

Capsules

subovoid, 2.5–3 × 3.5–4 mm, 3-lobed;

cocci rounded, smooth or puncticulate, glabrous;

columella 1.9–2.3 mm.

oblate, 1.7–2 × 2.2–2.7 mm, strigillose;

columella 1.5–1.9 mm.

Seeds

cream and brown mottled, oblong-ovoid to nearly globose, 2–2.5 × 1.4–1.7 mm, usually irregularly vermiculate-ridged and large-pitted, occasionally tuberculate or nearly smooth;

caruncle reniform, conic, 0.5–0.6 × 0.5–0.7 mm.

blackish brown, broadly ovoid, 3-angled in cross section, 1.2–1.4 × 1–1.1 mm, with 2 well-defined transverse ridges that do not pass through abaxial keel.

Cyathial

arrangement: terminal pleiochasial branches 3, each 2-branched;

pleiochasial bracts obovate to orbiculate-reniform, wider than distal leaves;

dichasial bracts usually connate 1/3–1/2 length (often only on one side), rarely only connate basally, triangular ovate to reniform, base truncate to perfoliate, margins erose-denticulate to subentire, apex rounded to obtuse, rarely apiculate;

axillary cymose branches 0–5.

Cyathia

peduncle 0–0.5 mm.

solitary at distal nodes;

peduncle 0.5–1.6 mm.

2n

= 14.

Euphorbia crenulata

Euphorbia rayturneri

Phenology Flowering and fruiting spring–summer. Flowering and fruiting late summer–early fall.
Habitat Conifer, oak, and mixed forests, coastal scrub, grasslands, barrens and outcrops, roadsides. Desert grasslands.
Elevation 30–1800 m. (100–5900 ft.) 1400–1700 m. (4600–5600 ft.)
Distribution
from FNA
CA; CO; NM; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
NM
Discussion

Euphorbia crenulata is most common in the central valleys of California and southern Oregon; it occurs disjunctly in southwestern Colorado and northwestern New Mexico. Previous reports from Arizona are based on misidentified specimens. Euphorbia crenulata is closely related to E. commutata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia rayturneri is known from only three collections in extreme southwestern New Mexico. Given its close proximity to the Mexican border, the species may also occur in the adjacent states of Chihuahua or Sonora.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 301. FNA vol. 12, p. 286.
Parent taxa Euphorbiaceae > Euphorbia > subg. Esula Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Tithymalus crenulatus
Name authority Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) V. W. Steinmann & Jercinovic: Novon 22: 482, figs. 1, 2. (2013)
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