Euphorbia crenulata |
Euphorbia rayturneri |
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Chinese caps, western wood spurge |
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Habit | Herbs, usually biennial, occasionally annual, with taproot. | Herbs, annual, with slender taproot. |
Stems | erect, sometimes decumbent at base, unbranched or branched, 12–40 cm, glabrous. |
prostrate, 4–8 cm, uniformly strigillose. |
Leaves | petiole 0–2 mm; blade obovate-spatulate to oblanceolate, 8–22 × 3–10 mm, base broadly attenuate, margins entire or slightly crisped, apex obtuse to ± rounded, minutely apiculate, surfaces glabrous; venation pinnate, midvein prominent. |
opposite; stipules distinct, filiform, 0.6–0.9 mm, pilose; petiole 0.5–0.9 mm, strigillose; blade ovate to elliptic, often slightly falcate, 5–11 × 2–5 mm, base asymmetric, one side cordate, other round to attenuate, margins sharply serrulate, apex acute, surfaces often with red spot toward middle, abaxial surface strigillose, adaxial surface glabrescent; only midvein conspicuous. |
Involucre | campanulate, 1.8–2.1 × 1.6–1.8 mm, glabrous; glands 4, crescent-shaped, 0.6–1.2 × 1.5–2.3 mm; horns slightly divergent to slightly convergent, 0.4–0.6 mm. |
obconic, 0.9–1.2 × 0.8–1.2 mm, strigillose; glands 4, green, yellow, or light pink, circular to oblong, 0.2 × 0.2–0.3 mm; appendages absent or green, yellow, or light pink, forming narrow margin on distal portion of gland, 0–0.1 × 0.2–0.3 mm, distal margin entire. |
Staminate flowers | 11–18. |
5–8. |
Pistillate flowers | ovary glabrous; styles 0.9–1.4 mm, 2-fid. |
ovary strigillose-canescent; styles 0.3–0.4 mm, unbranched. |
Capsules | subovoid, 2.5–3 × 3.5–4 mm, 3-lobed; cocci rounded, smooth or puncticulate, glabrous; columella 1.9–2.3 mm. |
oblate, 1.7–2 × 2.2–2.7 mm, strigillose; columella 1.5–1.9 mm. |
Seeds | cream and brown mottled, oblong-ovoid to nearly globose, 2–2.5 × 1.4–1.7 mm, usually irregularly vermiculate-ridged and large-pitted, occasionally tuberculate or nearly smooth; caruncle reniform, conic, 0.5–0.6 × 0.5–0.7 mm. |
blackish brown, broadly ovoid, 3-angled in cross section, 1.2–1.4 × 1–1.1 mm, with 2 well-defined transverse ridges that do not pass through abaxial keel. |
Cyathial | arrangement: terminal pleiochasial branches 3, each 2-branched; pleiochasial bracts obovate to orbiculate-reniform, wider than distal leaves; dichasial bracts usually connate 1/3–1/2 length (often only on one side), rarely only connate basally, triangular ovate to reniform, base truncate to perfoliate, margins erose-denticulate to subentire, apex rounded to obtuse, rarely apiculate; axillary cymose branches 0–5. |
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Cyathia | peduncle 0–0.5 mm. |
solitary at distal nodes; peduncle 0.5–1.6 mm. |
2n | = 14. |
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Euphorbia crenulata |
Euphorbia rayturneri |
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Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting late summer–early fall. |
Habitat | Conifer, oak, and mixed forests, coastal scrub, grasslands, barrens and outcrops, roadsides. | Desert grasslands. |
Elevation | 30–1800 m. (100–5900 ft.) | 1400–1700 m. (4600–5600 ft.) |
Distribution |
CA; CO; NM; OR
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NM |
Discussion | Euphorbia crenulata is most common in the central valleys of California and southern Oregon; it occurs disjunctly in southwestern Colorado and northwestern New Mexico. Previous reports from Arizona are based on misidentified specimens. Euphorbia crenulata is closely related to E. commutata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia rayturneri is known from only three collections in extreme southwestern New Mexico. Given its close proximity to the Mexican border, the species may also occur in the adjacent states of Chihuahua or Sonora. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 301. | FNA vol. 12, p. 286. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Tithymalus crenulatus | |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) | V. W. Steinmann & Jercinovic: Novon 22: 482, figs. 1, 2. (2013) |
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