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Chinese caps, western wood spurge

blue weed, ground spurge, prostrate sandmat, prostrate spurge or sandmat

Habit Herbs, usually biennial, occasionally annual, with taproot. Herbs, annual, with taproot.
Stems

erect, sometimes decumbent at base, unbranched or branched, 12–40 cm, glabrous.

prostrate to decumbent, usually not mat-forming, 10–30 cm, crisped-villous to glabrate (proximally).

Leaves

petiole 0–2 mm;

blade obovate-spatulate to oblanceolate, 8–22 × 3–10 mm, base broadly attenuate, margins entire or slightly crisped, apex obtuse to ± rounded, minutely apiculate, surfaces glabrous;

venation pinnate, midvein prominent.

opposite;

stipules connate (lower side) or distinct (upper side), triangular-subulate, often lacerate distally, 0.5–1 mm, short crisped-villous to glabrate;

petiole 0.5–1.5 mm, usually short crisped-villous, sometimes wooly at distal nodes;

blade broadly elliptic to elliptic-oblong, ovate-spatulate, or ovate, 3–11(–15) × 3–6(–8) mm, base slightly asymmetric, rounded to slightly cordate and oblique, margins serrulate at least in distal 1/2, sometimes obscurely so, apex obtuse, abaxial surface finely crisped-villous, adaxial surface usually glabrous or glabrate, sometimes sparsely crisped-villous; 3-veined from base.

Involucre

campanulate, 1.8–2.1 × 1.6–1.8 mm, glabrous;

glands 4, crescent-shaped, 0.6–1.2 × 1.5–2.3 mm;

horns slightly divergent to slightly convergent, 0.4–0.6 mm.

obconic, 0.6–0.9 × 0.5 mm, crisped-villous or glabrous;

glands 4, reddish, oval to oblong, 0.1 × 0.1–0.2 mm;

appendages white to pink, rudimentary, 0–0.2 mm, distal margin entire or irregularly scalloped.

Staminate flowers

11–18.

3–6.

Pistillate flowers

ovary glabrous;

styles 0.9–1.4 mm, 2-fid.

ovary densely crisped-villous;

styles 0.1 mm, 2-fid nearly entire length.

Capsules

subovoid, 2.5–3 × 3.5–4 mm, 3-lobed;

cocci rounded, smooth or puncticulate, glabrous;

columella 1.9–2.3 mm.

broadly ovoid, 1.2–2 × 1.4–1.5 mm, crisped-villous along keels and toward base, often glabrous between keels;

columella 1–1.2 mm.

Seeds

cream and brown mottled, oblong-ovoid to nearly globose, 2–2.5 × 1.4–1.7 mm, usually irregularly vermiculate-ridged and large-pitted, occasionally tuberculate or nearly smooth;

caruncle reniform, conic, 0.5–0.6 × 0.5–0.7 mm.

white but with barely concealed brown surface beneath, ovoid, sharply 4-angled in cross section, abaxial faces plane to convex, adaxial faces concave, 0.8–1.1 × 0.5–0.7 mm, with several narrow, sharp, slightly irregular, transverse ridges.

Cyathial

arrangement: terminal pleiochasial branches 3, each 2-branched;

pleiochasial bracts obovate to orbiculate-reniform, wider than distal leaves;

dichasial bracts usually connate 1/3–1/2 length (often only on one side), rarely only connate basally, triangular ovate to reniform, base truncate to perfoliate, margins erose-denticulate to subentire, apex rounded to obtuse, rarely apiculate;

axillary cymose branches 0–5.

Cyathia

peduncle 0–0.5 mm.

solitary or in small, cymose clusters at distal nodes or on congested, axillary branches;

peduncle 1–2 mm.

2n

= 18.

Euphorbia crenulata

Euphorbia prostrata

Phenology Flowering and fruiting spring–summer. Flowering and fruiting early spring–fall.
Habitat Conifer, oak, and mixed forests, coastal scrub, grasslands, barrens and outcrops, roadsides. Disturbed areas, fields, gardens, sidewalks, sandy places, ballast piles.
Elevation 30–1800 m. (100–5900 ft.) 0–1400 m. (0–4600 ft.)
Distribution
from FNA
CA; CO; NM; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; AZ; CA; CO; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; NC; NE; NM; OH; OK; PA; SC; SD; TN; TX; UT; VA; WV; WY; Mexico; Central America; South America; West Indies [Introduced in Eurasia, Africa, Pacific Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia crenulata is most common in the central valleys of California and southern Oregon; it occurs disjunctly in southwestern Colorado and northwestern New Mexico. Previous reports from Arizona are based on misidentified specimens. Euphorbia crenulata is closely related to E. commutata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia prostrata is native to tropical America and possibly into the southern part of the flora area. It is likely adventive throughout most of the northern part of its range. It is widely naturalized throughout much of the rest of the tropics.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 301. FNA vol. 12, p. 285.
Parent taxa Euphorbiaceae > Euphorbia > subg. Esula Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Tithymalus crenulatus Chamaesyce prostrata
Name authority Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) Aiton: Hort. Kew. 2: 139. (1789)
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