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Chinese caps, western wood spurge

Habit Herbs, usually biennial, occasionally annual, with taproot. Herbs, annual, with taproot.
Stems

erect, sometimes decumbent at base, unbranched or branched, 12–40 cm, glabrous.

prostrate to ascending, drooping at tips, 15–30 cm, glabrous.

Leaves

petiole 0–2 mm;

blade obovate-spatulate to oblanceolate, 8–22 × 3–10 mm, base broadly attenuate, margins entire or slightly crisped, apex obtuse to ± rounded, minutely apiculate, surfaces glabrous;

venation pinnate, midvein prominent.

opposite;

stipules distinct, divided nearly to base into linear-filiform segments, 1–2 mm, glabrous;

petiole 0.5–1.3 mm, glabrous;

blade narrowly oblong to oblong-obovate, often ± falcate, 5–16 × 1–4.5 mm, base subsymmetric to strongly oblique, margins sparsely spinulose-serrulate, apex rounded or broadly acute, abaxial surface pale grayish green, adaxial surface sometimes with reddish streak along midvein, both surfaces not papillate, glabrous; 3–5-veined at base.

Involucre

campanulate, 1.8–2.1 × 1.6–1.8 mm, glabrous;

glands 4, crescent-shaped, 0.6–1.2 × 1.5–2.3 mm;

horns slightly divergent to slightly convergent, 0.4–0.6 mm.

obconic, 0.7–0.9 × 0.4–0.6 mm, glabrous;

glands 4, green to yellow-green, subequal, oblong, 0.1–0.2 × 0.2–0.3 mm;

appendages white to reddish tinged, lunate to oblong, 0.1–0.3 × 0.3–0.5 mm (2 ± 2 times longer than other 2), entire or coarsely toothed.

Staminate flowers

11–18.

2–5.

Pistillate flowers

ovary glabrous;

styles 0.9–1.4 mm, 2-fid.

ovary glabrous;

styles 0.2–0.3 mm, 2-fid at apex to nearly 1/2 length.

Capsules

subovoid, 2.5–3 × 3.5–4 mm, 3-lobed;

cocci rounded, smooth or puncticulate, glabrous;

columella 1.9–2.3 mm.

ovoid, cocci not elongated nor terminating in empty portion, 1.5 × 1.7 mm, glabrous;

columella 1.3 mm.

Seeds

cream and brown mottled, oblong-ovoid to nearly globose, 2–2.5 × 1.4–1.7 mm, usually irregularly vermiculate-ridged and large-pitted, occasionally tuberculate or nearly smooth;

caruncle reniform, conic, 0.5–0.6 × 0.5–0.7 mm.

reddish brown to brown, pyramidal to oblong-ovoid, weakly 4-angled in cross section, 0.9–1 × 0.7 mm, minutely beaded, with 3–4 broad, rounded, transverse ridges that do not interrupt abaxial keel.

Cyathial

arrangement: terminal pleiochasial branches 3, each 2-branched;

pleiochasial bracts obovate to orbiculate-reniform, wider than distal leaves;

dichasial bracts usually connate 1/3–1/2 length (often only on one side), rarely only connate basally, triangular ovate to reniform, base truncate to perfoliate, margins erose-denticulate to subentire, apex rounded to obtuse, rarely apiculate;

axillary cymose branches 0–5.

Cyathia

peduncle 0–0.5 mm.

solitary at distal nodes or on congested, axillary branches;

peduncle 0.1–0.4 mm.

Euphorbia crenulata

Euphorbia meganaesos

Phenology Flowering and fruiting spring–summer. Flowering and fruiting late spring–late summer.
Habitat Conifer, oak, and mixed forests, coastal scrub, grasslands, barrens and outcrops, roadsides. Sandy beaches, edges of marshes, coastal prairies, roadsides.
Elevation 30–1800 m. (100–5900 ft.) 0–10 m. (0–0 ft.)
Distribution
from FNA
CA; CO; NM; OR
[WildflowerSearch map]
[BONAP county map]
from FNA
LA; TX
Discussion

Euphorbia crenulata is most common in the central valleys of California and southern Oregon; it occurs disjunctly in southwestern Colorado and northwestern New Mexico. Previous reports from Arizona are based on misidentified specimens. Euphorbia crenulata is closely related to E. commutata.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia meganaesos is known only from coastal areas of southern Louisiana and adjacent Texas. This species was often considered conspecific with E. maculata in the past, but it differs from that species in being entirely glabrous.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 301. FNA vol. 12, p. 276.
Parent taxa Euphorbiaceae > Euphorbia > subg. Esula Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Tithymalus crenulatus
Name authority Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) Featherman: Rep. (Annual) Board Supervisors Louisiana State Seminary Learning Military Acad. 1870: 71, 105. (1871)
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