Euphorbia crenulata |
Euphorbia deltoidea |
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Chinese caps, western wood spurge |
wedge sandmat |
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Habit | Herbs, usually biennial, occasionally annual, with taproot. | Herbs, perennial, delicate, with woody, thickened taproot, 15 mm diam.. | ||||||||||||||||
Stems | erect, sometimes decumbent at base, unbranched or branched, 12–40 cm, glabrous. |
prostrate, ascending, or erect, often numerous and wiry, less than 0.1 mm diam., 5–20 cm, glabrous, puberulent, canescent, villous, or hirsute, shorter hairs often uncinate and longer hairs straight or irregularly twisted. |
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Leaves | petiole 0–2 mm; blade obovate-spatulate to oblanceolate, 8–22 × 3–10 mm, base broadly attenuate, margins entire or slightly crisped, apex obtuse to ± rounded, minutely apiculate, surfaces glabrous; venation pinnate, midvein prominent. |
opposite; stipules distinct, triangular, sometimes lacerate or ciliate, 0.2–0.3 mm, glabrous or hairy; petiole 0.3–1 mm, glabrous or hairy; blade narrowly to broadly deltate, cordate, or reniform, 2–5(–7) × 1–4.5(–5) mm, base asymmetric, cordate to rounded, margins entire, ± revolute, apex obtuse or rounded, surfaces glabrous or hairy; only midvein conspicuous. |
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Involucre | campanulate, 1.8–2.1 × 1.6–1.8 mm, glabrous; glands 4, crescent-shaped, 0.6–1.2 × 1.5–2.3 mm; horns slightly divergent to slightly convergent, 0.4–0.6 mm. |
turbinate to campanulate, 0.8–1 × 1.1–1.3 mm, glabrous or hairy; glands 4, green to yellow-green, oblong to subcircular, 0.2–0.4 × 0.4–0.6 mm; appendages absent or white, forming narrow rim at edge of gland, rarely slightly wider than gland, (0–)0.1(–0.3) × 0.4–0.6 mm, distal margin entire. |
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Staminate flowers | 11–18. |
8–14. |
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Pistillate flowers | ovary glabrous; styles 0.9–1.4 mm, 2-fid. |
ovary glabrous or hairy, subtended by triangular pad of tissue; styles spreading, 0.3–0.4 mm, 2-fid 1/2 to nearly entire length. |
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Capsules | subovoid, 2.5–3 × 3.5–4 mm, 3-lobed; cocci rounded, smooth or puncticulate, glabrous; columella 1.9–2.3 mm. |
broadly deltoid, 1.2–1.5 × 2–2.2 mm, glabrous or hairy; columella 0.9–1.3 mm. |
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Seeds | cream and brown mottled, oblong-ovoid to nearly globose, 2–2.5 × 1.4–1.7 mm, usually irregularly vermiculate-ridged and large-pitted, occasionally tuberculate or nearly smooth; caruncle reniform, conic, 0.5–0.6 × 0.5–0.7 mm. |
reddish brown, ovoid, 4-angled in cross section, 0.8–1.2 × 0.5–0.6 mm, obscurely wrinkled. |
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Cyathial | arrangement: terminal pleiochasial branches 3, each 2-branched; pleiochasial bracts obovate to orbiculate-reniform, wider than distal leaves; dichasial bracts usually connate 1/3–1/2 length (often only on one side), rarely only connate basally, triangular ovate to reniform, base truncate to perfoliate, margins erose-denticulate to subentire, apex rounded to obtuse, rarely apiculate; axillary cymose branches 0–5. |
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Cyathia | peduncle 0–0.5 mm. |
solitary at distal nodes; peduncle 0.7–1.5 mm. |
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Euphorbia crenulata |
Euphorbia deltoidea |
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Phenology | Flowering and fruiting spring–summer. | |||||||||||||||||
Habitat | Conifer, oak, and mixed forests, coastal scrub, grasslands, barrens and outcrops, roadsides. | |||||||||||||||||
Elevation | 30–1800 m. (100–5900 ft.) | |||||||||||||||||
Distribution |
CA; CO; NM; OR
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FL |
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Discussion | Euphorbia crenulata is most common in the central valleys of California and southern Oregon; it occurs disjunctly in southwestern Colorado and northwestern New Mexico. Previous reports from Arizona are based on misidentified specimens. Euphorbia crenulata is closely related to E. commutata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 4 (4 in the flora). Euphorbia deltoidea comprises four narrowly endemic subspecies, all of which are endangered due to habitat loss and fragmentation. The subspecies occur in pine rockland habitat that is free of shrubby undergrowth. Periodic fires are required to keep the rockland habitat open. Subspecies serpyllum is restricted to Big Pine Key, Monroe County, whereas the other subspecies are found only in Miami-Dade County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 301. | FNA vol. 12, p. 265. | ||||||||||||||||
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum | ||||||||||||||||
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Synonyms | Tithymalus crenulatus | Chamaesyce deltoidea | ||||||||||||||||
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) | Engelmann ex Chapman: Fl. South. U.S. ed. 2, 647. (1883) | ||||||||||||||||
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