Euphorbia crenulata |
Euphorbia chaetocalyx |
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Chinese caps, western wood spurge |
bristlecup sandmat |
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Habit | Herbs, usually biennial, occasionally annual, with taproot. | Herbs, perennial, with woody, thickened taproot. | ||||
Stems | erect, sometimes decumbent at base, unbranched or branched, 12–40 cm, glabrous. |
usually erect, rarely slightly decumbent, often densely clustered from top of woody crown, 3–15 cm, glabrous. |
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Leaves | petiole 0–2 mm; blade obovate-spatulate to oblanceolate, 8–22 × 3–10 mm, base broadly attenuate, margins entire or slightly crisped, apex obtuse to ± rounded, minutely apiculate, surfaces glabrous; venation pinnate, midvein prominent. |
opposite; stipules distinct, narrowly linear, usually entire, 0.5–1 mm, glabrous; petiole 0.5–1 mm, glabrous; blade ovate to lanceolate or oblong- or linear-lanceolate, 3–11 × 0.8–3(–5) mm, base slightly asymmetric, short-tapered, occasionally one side slightly rounded, margins entire, apex acute or short-acuminate, surfaces glabrous; only midvein conspicuous. |
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Involucre | campanulate, 1.8–2.1 × 1.6–1.8 mm, glabrous; glands 4, crescent-shaped, 0.6–1.2 × 1.5–2.3 mm; horns slightly divergent to slightly convergent, 0.4–0.6 mm. |
campanulate to turbinate, 0.8–1.4 × 0.8–1 mm, glabrous; glands 4, yellow-brown to reddish, concave or convex, elliptic or oval, 0.2–0.4 × 0.4–0.6 mm; appendages absent or white, lanceolate-deltate to straplike, 0.2–1.1 × 0.2–0.9 mm, distal margin entire, crenate, or deeply cleft or divided. |
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Staminate flowers | 11–18. |
25–35. |
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Pistillate flowers | ovary glabrous; styles 0.9–1.4 mm, 2-fid. |
ovary glabrous; styles 0.3–0.4 mm, 2-fid 1/2 length. |
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Capsules | subovoid, 2.5–3 × 3.5–4 mm, 3-lobed; cocci rounded, smooth or puncticulate, glabrous; columella 1.9–2.3 mm. |
depressed-ovoid to depressed-globose, 1.7–2.1 × 1.6–2.4 mm, glabrous; columella 1.2–1.8 mm. |
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Seeds | cream and brown mottled, oblong-ovoid to nearly globose, 2–2.5 × 1.4–1.7 mm, usually irregularly vermiculate-ridged and large-pitted, occasionally tuberculate or nearly smooth; caruncle reniform, conic, 0.5–0.6 × 0.5–0.7 mm. |
white, ovoid-pyramidal, prominently 4-angled in cross section, 1.6–2 × 1–1.2 mm, smooth to slightly wrinkled. |
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Cyathial | arrangement: terminal pleiochasial branches 3, each 2-branched; pleiochasial bracts obovate to orbiculate-reniform, wider than distal leaves; dichasial bracts usually connate 1/3–1/2 length (often only on one side), rarely only connate basally, triangular ovate to reniform, base truncate to perfoliate, margins erose-denticulate to subentire, apex rounded to obtuse, rarely apiculate; axillary cymose branches 0–5. |
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Cyathia | peduncle 0–0.5 mm. |
solitary at distal nodes; peduncle 0.8–1.3 mm. |
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Euphorbia crenulata |
Euphorbia chaetocalyx |
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Phenology | Flowering and fruiting spring–summer. | |||||
Habitat | Conifer, oak, and mixed forests, coastal scrub, grasslands, barrens and outcrops, roadsides. | |||||
Elevation | 30–1800 m. (100–5900 ft.) | |||||
Distribution |
CA; CO; NM; OR
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AZ; NM; TX; n Mexico
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Discussion | Euphorbia crenulata is most common in the central valleys of California and southern Oregon; it occurs disjunctly in southwestern Colorado and northwestern New Mexico. Previous reports from Arizona are based on misidentified specimens. Euphorbia crenulata is closely related to E. commutata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Varieties 2 (2 in the flora). Euphorbia chaetocalyx is similar to E. fendleri but can generally be distinguished from that species by its narrow, acute leaves and ± erect stems. Some authors have used the presence or absence and shape of the involucral gland appendages to help separate E. chaetocalyx from E. fendleri, but those characters appear highly variable and of little taxonomic utility. Some individuals from western Texas (Culberson and El Paso counties) and southern New Mexico appear intermediate with E. fendleri. The specific epithet of E. chaetocalyx refers to the bristly perianthlike segments that subtend the ovary, but these structures are found intermittently in both E. chaetocalyx and E. fendleri. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 301. | FNA vol. 12, p. 263. | ||||
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | Tithymalus crenulatus | E. fendleri var. chaetocalyx, Chamaesyce chaetocalyx | ||||
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) | (Boissier) Tidestrom: Proc. Biol. Soc. Wash. 48: 40. (1935) | ||||
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