Euphorbia crenulata |
Euphorbia astyla |
|
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Chinese caps, western wood spurge |
alkali spurge, Pecos spurge |
|
Habit | Herbs, usually biennial, occasionally annual, with taproot. | Herbs, perennial, with woody or fibrous-fleshy taproot, 5–12 mm thick. |
Stems | erect, sometimes decumbent at base, unbranched or branched, 12–40 cm, glabrous. |
decumbent, ascending, or erect, few to many emerging from woody crown, 5–25(–50) cm, glabrous. |
Leaves | petiole 0–2 mm; blade obovate-spatulate to oblanceolate, 8–22 × 3–10 mm, base broadly attenuate, margins entire or slightly crisped, apex obtuse to ± rounded, minutely apiculate, surfaces glabrous; venation pinnate, midvein prominent. |
opposite; stipules connate into deltate scale, 0.2–0.5 mm, minutely lacerate at apex, glabrous; petiole 0–0.2(–0.3) mm, glabrous; blade orbiculate-reniform to acute-cordate, 2–5(–8) × 2–5(–6) mm, base ± asymmetric, cordate to auriculate, sometimes clasping stem, margins entire, apex narrowly acute, surfaces glabrous; 2- or 3-veined from base, but usually only midvein conspicuous. |
Involucre | campanulate, 1.8–2.1 × 1.6–1.8 mm, glabrous; glands 4, crescent-shaped, 0.6–1.2 × 1.5–2.3 mm; horns slightly divergent to slightly convergent, 0.4–0.6 mm. |
broadly campanulate, 0.8–1.4 × 0.9–1.4 mm, glabrous; glands 4, yellow-green to brownish, oblong, 0.2–0.3 × 0.5–0.7 mm; appendages white, flabellate to oblong, 0.1–0.2(–0.5) × 0.4–0.8 mm, distal margin entire or dentate-crenate. |
Staminate flowers | 11–18. |
22–26. |
Pistillate flowers | ovary glabrous; styles 0.9–1.4 mm, 2-fid. |
ovary glabrous; styles 0.3–0.4 mm, unbranched, thickened-clavate. |
Capsules | subovoid, 2.5–3 × 3.5–4 mm, 3-lobed; cocci rounded, smooth or puncticulate, glabrous; columella 1.9–2.3 mm. |
ovoid and broadly triangular, 1.5–1.9(–2.5) × 1.4–1.6(–2.2) mm, glabrous; columella 1.2–1.8 mm. |
Seeds | cream and brown mottled, oblong-ovoid to nearly globose, 2–2.5 × 1.4–1.7 mm, usually irregularly vermiculate-ridged and large-pitted, occasionally tuberculate or nearly smooth; caruncle reniform, conic, 0.5–0.6 × 0.5–0.7 mm. |
white, oblong, 4-angled in cross section, adaxial faces slightly concave, with long raphe between, 1.5–1.8 × 0.7–1 mm, markedly foveolate, with irregular to ± parallel or anastomosing ridges. |
Cyathial | arrangement: terminal pleiochasial branches 3, each 2-branched; pleiochasial bracts obovate to orbiculate-reniform, wider than distal leaves; dichasial bracts usually connate 1/3–1/2 length (often only on one side), rarely only connate basally, triangular ovate to reniform, base truncate to perfoliate, margins erose-denticulate to subentire, apex rounded to obtuse, rarely apiculate; axillary cymose branches 0–5. |
|
Cyathia | peduncle 0–0.5 mm. |
solitary at distal nodes; peduncle 0.3–1(–1.5) mm. |
2n | = 28. |
|
Euphorbia crenulata |
Euphorbia astyla |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting late spring–early fall. |
Habitat | Conifer, oak, and mixed forests, coastal scrub, grasslands, barrens and outcrops, roadsides. | Desert, grasslands, limestone substrates, usually on very saline or alkaline soils. |
Elevation | 30–1800 m. (100–5900 ft.) | 700–1100 m. (2300–3600 ft.) |
Distribution |
CA; CO; NM; OR
|
TX; Mexico (Coahuila, Nuevo León) |
Discussion | Euphorbia crenulata is most common in the central valleys of California and southern Oregon; it occurs disjunctly in southwestern Colorado and northwestern New Mexico. Previous reports from Arizona are based on misidentified specimens. Euphorbia crenulata is closely related to E. commutata. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia astyla is a specialist on halophytic, alkaline soils and is known in the flora area only in part of Pecos County. The species is closely related to E. jejuna but differs in its sessile or sub-sessile leaves with a cordate-auriculate base and involucral gland appendages that are not deeply lobed or cleft. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 301. | FNA vol. 12, p. 261. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Tithymalus crenulatus | Chamaesyce astyla |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) | Engelmann ex Boissier: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 40. (1862) |
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