Euphorbia cordifolia |
Euphorbia exstipulata |
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heartleaf sandmat |
square-seed spurge, squareseed or Clark Mountain spurge |
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Habit | Herbs, annual, with taproot. | Herbs, annual, with slender taproot. |
Stems | prostrate, occasionally mat-forming, 10–43 cm, glabrous. |
erect, 5–26 cm, uniformly puberulent to hispidulous or glabrous; branches arcuate. |
Leaves | opposite; stipules usually distinct, occasionally connate at base, filiform, 1–1.2(–2.8) mm, usually glabrous, rarely pilose; petiole 0.4–1 mm, usually glabrous; blade ovate to oblong, 4.4–12 × 2.6–7.6 mm, base asymmetric, cordate to rounded, margins entire, apex rounded to mucronulate, surfaces glabrous; only midvein conspicuous. |
opposite; petiole 1–3 mm, often indistinct, glabrous or puberulent; blade linear to narrowly elliptic or ovate, 14–42 × 3–28 mm, base attenuate, margins coarsely serrate, occasionally revolute, apex acute to obtuse, abaxial surface sparsely hispidulous to strigillose, adaxial surface glabrous; midvein conspicuous. |
Involucre | campanulate, 1–1.3 × 1–1.3 mm, glabrous; glands 4, yellowish to pink, elliptic, 0.3–0.5 × 0.5–1 mm; appendages whitish to pink, sometimes drying red, elliptic to ovoid, 1.1–1.5 × 1.2–1.9 mm, distal margin entire, retuse, or erose. |
turbinate to campanulate, 1.1–1.5 × 1–1.3 mm, glabrous, pilose or puberulent; involucral lobes divided into several linear lobes; glands 4(–5), yellow to pink, sessile and broadly attached, 0.2 × 0.3–0.4 mm, opening oblong to nearly circular, glabrous; appendages usually petaloid, white to pink, ovate to trapezoidal, occasionally absent, not incurved and covering glands, 0.2–0.4 × 0.3–0.8 mm, entire, undulate, or conspicuously divided into triangular segments, glabrous. |
Staminate flowers | 5–40. |
10–12. |
Pistillate flowers | ovary glabrous; styles 0.5–0.8 mm, 2-fid nearly entire length. |
ovary puberulent on keels, styles 0.8–1.1 mm, 2-fid 1/2 to nearly entire length. |
Capsules | ovoid, 2–3 mm diam., glabrous; columella 1.2–2.7 mm. |
broadly depressed-oblong to ovoid, 2.7–3.3 × 3.1–3.9 mm, puberulent (with appressed hairs usually concentrated on keels); columella 1.9–2.5 mm. |
Seeds | gray or tan with dark brown mottling, ovoid, bluntly 3–4-angled in cross section, 1.8–2.1 × 1.2–1.4 mm, smooth to rugose. |
white to gray or light brown, ovoid, bluntly 4-angled in cross section, 1.9–2.5 × 1.4–1.7 mm, tuberculate, often with 2 transverse ridges; caruncle 0.1 × 0.2 mm. |
Cyathia | solitary at distal nodes; peduncle 0.9–3 mm. |
peduncle 1–1.9 mm. |
Cyathial | arrangement: terminal cymose or dichasial branches usually 1–2, occasionally reduced to monochasia, 1–2-branched; pleiochasial bracts 2–4, often whorled, wholly green or paler green at base, similar in shape and size to distal leaves or slightly narrower; dichasial bracts similar in shape to distal leaves but smaller or highly reduced. |
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Euphorbia cordifolia |
Euphorbia exstipulata |
|
Phenology | Flowering and fruiting summer–fall. | Flowering and fruiting summer–fall. |
Habitat | Xeric oak-pine scrub, pine-barrens, sand barrens, sandy stream banks. | Desert scrub, grasslands, mesquite savannas, oak and oak-juniper woodlands. |
Elevation | 0–200 m. (0–700 ft.) | 800–2000 m. (2600–6600 ft.) |
Distribution |
AL; AR; FL; GA; LA; MS; NC; OK; SC; TX
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AZ; CA; NM; TX; UT; WY; Mexico (Chihuahua, Coahuila, Durango, Guanajuato, San Luis Potosí, Sonora, Zacatecas)
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Discussion | Euphorbia cordifolia is easily identified by its cordate to rounded leaf base and distinctive filiform stipules. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia exstipulata is native from Texas to California and northern Mexico. The species was found once in the late nineteenth century in Wyoming but has not been re-collected there. Broad-leaved plants have been segregated as var. lata, but the variation in leaf shape is continuous and no varieties are formally recognized here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 265. | FNA vol. 12, p. 322. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Poinsettia |
Sibling taxa | ||
Synonyms | Chamaesyce cordifolia | E. exstipulata var. lata |
Name authority | Elliott: Sketch. Bot. S. Carolina 2: 656. (1824) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) |
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