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heartleaf sandmat

Habit Herbs, annual, with taproot.
Stems

prostrate, occasionally mat-forming, 10–43 cm, glabrous.

Leaves

opposite;

stipules usually distinct, occasionally connate at base, filiform, 1–1.2(–2.8) mm, usually glabrous, rarely pilose;

petiole 0.4–1 mm, usually glabrous;

blade ovate to oblong, 4.4–12 × 2.6–7.6 mm, base asymmetric, cordate to rounded, margins entire, apex rounded to mucronulate, surfaces glabrous;

only midvein conspicuous.

Involucre

campanulate, 1–1.3 × 1–1.3 mm, glabrous;

glands 4, yellowish to pink, elliptic, 0.3–0.5 × 0.5–1 mm;

appendages whitish to pink, sometimes drying red, elliptic to ovoid, 1.1–1.5 × 1.2–1.9 mm, distal margin entire, retuse, or erose.

Staminate flowers

5–40.

Pistillate flowers

ovary glabrous;

styles 0.5–0.8 mm, 2-fid nearly entire length.

Capsules

ovoid, 2–3 mm diam., glabrous;

columella 1.2–2.7 mm.

Seeds

gray or tan with dark brown mottling, ovoid, bluntly 3–4-angled in cross section, 1.8–2.1 × 1.2–1.4 mm, smooth to rugose.

Cyathia

solitary at distal nodes;

peduncle 0.9–3 mm.

Euphorbia cordifolia

Phenology Flowering and fruiting summer–fall.
Habitat Xeric oak-pine scrub, pine-barrens, sand barrens, sandy stream banks.
Elevation 0–200 m. (0–700 ft.)
Distribution
from FNA
AL; AR; FL; GA; LA; MS; NC; OK; SC; TX
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia cordifolia is easily identified by its cordate to rounded leaf base and distinctive filiform stipules.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 265.
Parent taxa Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce cordifolia
Name authority Elliott: Sketch. Bot. S. Carolina 2: 656. (1824)
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