Euphorbia chamaesula |
Euphorbia trachysperma |
|
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mountain spurge |
San Pedro River sandmat |
|
Habit | Herbs, perennial, with thick rootstock. | Herbs, annual, with slender taproot. |
Stems | erect, branched, 40–90 cm, glabrous. |
erect to ascending, 10–55 cm, glabrous. |
Leaves | petiole 0.5–1 mm; blade elliptic to oblong, 8–20(–40) × 3–6 mm, base truncate, rounded, or attenuate, margins entire, apex obtuse or acute, sometimes slightly mucronate, surfaces glabrous; venation inconspicuous, only midvein prominent. |
opposite; stipules distinct, narrowly triangular and often divided into 3–5 subulate to filiform segments, 0.8–1.9 mm, glabrous; petiole 1.2–3.1 mm, glabrous; blade oblong, elliptic, or ovate, 12–43 × 3–14 mm, base subsymmetric to asymmetric, rounded, attenuate, or with one side hemicordate and other side rounded, margins usually serrulate, rarely entire on some leaves or portion of blade, apex obtuse to acute, surfaces often with red blotch in center, glabrous; pinnately veined, often only midvein conspicuous. |
Involucre | turbinate or campanulate, 1.5–2.5 × 1.1–1.9 mm, glabrous; glands 4, crescent-shaped to semicircular, 0.5–0.8 × 1–1.8 mm; horns usually convergent, 0.2–0.8 mm. |
oblong, campanulate, or funnel-shaped, 1.5–2.6 × 1.1–2.3 mm, glabrous; glands 4, green to yellowish or tinged with red, subcircular to oblong, 0.4–0.7 × 0.5–1 mm; appendages absent or white, often forming narrow rim or oblong to ovate, 0–0.8 × 1.2 mm, distal margin entire. |
Staminate flowers | 8–12. |
50–60. |
Pistillate flowers | ovary glabrous; styles 1–1.8 mm, 2-fid. |
ovary glabrous; styles 0.5–0.6 mm, 2-fid 1/2 length. |
Capsules | depressed-ovoid, 4.3–5 × 5–6 mm, 3-lobed; cocci rounded, smooth, glabrous; columella 3.5–4 mm. |
oblate, 2.4–3.2 × 3–3.5 mm, glabrous; columella 2.3–2.5 mm. |
Seeds | gray to dark brown, ovoid-oblong, truncate at both ends, 2.6–3.4 × 2–2.6 mm, shallowly pitted to almost smooth; caruncle conic, 1 × 0.8 mm. |
light brown, ovoid, 4-angled in cross section, abaxial keel well developed, 1.9–2.3 × 1.3–1.4 mm, finely dimpled and papillate. |
Cyathial | arrangement: terminal pleiochasial branches 3–5(–6), each 3–4 times 2-branched; pleiochasial bracts ovate-lanceolate to slightly subpandurate, similar in size or wider than distal leaves; dichasial bracts distinct, lanceolate to broadly ovate, base usually truncate to rounded or subcordate, sometimes attenuate, margins entire or slightly crenulate, apex usually obtuse to acute, occasionally acuminate; axillary cymose branches 2–8. |
|
Cyathia | peduncle 1–3 mm. |
solitary at distal nodes; peduncle 1.4–2.4 mm. |
2n | = 26. |
|
Euphorbia chamaesula |
Euphorbia trachysperma |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting late summer–early winter. |
Habitat | Clearings in ponderosa pine forests, montane roadsides, dry streambeds, creek banks, sandy and gravelly soils. | Desert scrub, desert grasslands, mesquite woodlands. |
Elevation | 1700–2700 m. (5600–8900 ft.) | 200–1200 m. (700–3900 ft.) |
Distribution |
AZ; NM; Mexico (Chihuahua, Sonora)
|
AZ; Mexico (Baja California Sur, Sonora) |
Discussion | Euphorbia chamaesula is easily distinguished from other perennial members of subg. Esula in western North America by its larger capsules and the vegetative shoots (without cyathia) that arise from the distal nodes of the stem proximal to the pleiochasia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia trachysperma may be expected in extreme southwestern New Mexico given the close proximity of some collections in southeastern Arizona. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 300. | FNA vol. 12, p. 291. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Tithymalus chamaesula | Chamaesyce trachysperma |
Name authority | Boissier: Cent. Euphorb., 38. (1860) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 189. (1859) |
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