Euphorbia chamaesula |
Euphorbia oblongata |
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mountain spurge |
Balkan spurge, egg-leaf spurge, oblong spurge |
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Habit | Herbs, perennial, with thick rootstock. | Herbs, perennial, with woody taproot. |
Stems | erect, branched, 40–90 cm, glabrous. |
erect, unbranched or densely branching, 80 cm, often densely villous (especially young stems and pleiochasial branches). |
Leaves | petiole 0.5–1 mm; blade elliptic to oblong, 8–20(–40) × 3–6 mm, base truncate, rounded, or attenuate, margins entire, apex obtuse or acute, sometimes slightly mucronate, surfaces glabrous; venation inconspicuous, only midvein prominent. |
petiole absent; blade oblong to narrowly obovate or lanceolate, 15–70 × 6–25 mm, base rounded or truncate, margins finely serrulate, apex obtuse, mucronulate, surfaces glabrous; venation inconspicuously pinnate, midvein prominent. |
Involucre | turbinate or campanulate, 1.5–2.5 × 1.1–1.9 mm, glabrous; glands 4, crescent-shaped to semicircular, 0.5–0.8 × 1–1.8 mm; horns usually convergent, 0.2–0.8 mm. |
cupulate to slightly turbinate, 1.5–2.5 × 1.3–1.5 mm, glabrous; glands 2–3, elliptic, 0.6–0.8 × 0.8–1.2 mm; horns absent. |
Staminate flowers | 8–12. |
15–40. |
Pistillate flowers | ovary glabrous; styles 1–1.8 mm, 2-fid. |
ovary glabrous; styles 1.5–2 mm, 2-fid. |
Capsules | depressed-ovoid, 4.3–5 × 5–6 mm, 3-lobed; cocci rounded, smooth, glabrous; columella 3.5–4 mm. |
globose, 3–4.5 × 3–4.5 mm, slightly 3-lobed; cocci rounded, verrucose-tuberculate, glabrous; columella 2.5–3.3 mm. |
Seeds | gray to dark brown, ovoid-oblong, truncate at both ends, 2.6–3.4 × 2–2.6 mm, shallowly pitted to almost smooth; caruncle conic, 1 × 0.8 mm. |
brown, ovoid, 2.4–2.6 × 1.3–2 mm, smooth, caruncle reniform, 0.2–0.3 × 0.8–0.9 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3–5(–6), each 3–4 times 2-branched; pleiochasial bracts ovate-lanceolate to slightly subpandurate, similar in size or wider than distal leaves; dichasial bracts distinct, lanceolate to broadly ovate, base usually truncate to rounded or subcordate, sometimes attenuate, margins entire or slightly crenulate, apex usually obtuse to acute, occasionally acuminate; axillary cymose branches 2–8. |
arrangement: terminal pleiochasial branches 3–5, each 2–3 times 2–4-branched; pleiochasial bracts ovate, similar in size to distal leaves; dichasial bracts distinct, ovate to suborbiculate, base truncate or rounded, margins entire or finely denticulate, apex obtuse, sometimes mucronulate; axillary cymose branches 0–4. |
Cyathia | peduncle 1–3 mm. |
peduncle 1–5 mm. |
2n | = 26. |
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Euphorbia chamaesula |
Euphorbia oblongata |
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Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting spring–fall. |
Habitat | Clearings in ponderosa pine forests, montane roadsides, dry streambeds, creek banks, sandy and gravelly soils. | Waste areas, disturbed sites, roadsides, fields, pastures. |
Elevation | 1700–2700 m. (5600–8900 ft.) | 30–900 m. (100–3000 ft.) |
Distribution |
AZ; NM; Mexico (Chihuahua, Sonora)
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CA; OR; WA; s Europe [Introduced in North America]
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Discussion | Euphorbia chamaesula is easily distinguished from other perennial members of subg. Esula in western North America by its larger capsules and the vegetative shoots (without cyathia) that arise from the distal nodes of the stem proximal to the pleiochasia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia oblongata is listed as a noxious weed by the states of California, Oregon, and Washington. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 300. | FNA vol. 12, p. 305. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Tithymalus chamaesula | Tithymalus oblongatus |
Name authority | Boissier: Cent. Euphorb., 38. (1860) | Grisebach: Spic. Fl. Rumel. 1: 136. (1843) |
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