Euphorbia chamaesula |
Euphorbia deltoidea |
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mountain spurge |
wedge sandmat |
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Habit | Herbs, perennial, with thick rootstock. | Herbs, perennial, delicate, with woody, thickened taproot, 15 mm diam.. | ||||||||||||||||
Stems | erect, branched, 40–90 cm, glabrous. |
prostrate, ascending, or erect, often numerous and wiry, less than 0.1 mm diam., 5–20 cm, glabrous, puberulent, canescent, villous, or hirsute, shorter hairs often uncinate and longer hairs straight or irregularly twisted. |
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Leaves | petiole 0.5–1 mm; blade elliptic to oblong, 8–20(–40) × 3–6 mm, base truncate, rounded, or attenuate, margins entire, apex obtuse or acute, sometimes slightly mucronate, surfaces glabrous; venation inconspicuous, only midvein prominent. |
opposite; stipules distinct, triangular, sometimes lacerate or ciliate, 0.2–0.3 mm, glabrous or hairy; petiole 0.3–1 mm, glabrous or hairy; blade narrowly to broadly deltate, cordate, or reniform, 2–5(–7) × 1–4.5(–5) mm, base asymmetric, cordate to rounded, margins entire, ± revolute, apex obtuse or rounded, surfaces glabrous or hairy; only midvein conspicuous. |
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Involucre | turbinate or campanulate, 1.5–2.5 × 1.1–1.9 mm, glabrous; glands 4, crescent-shaped to semicircular, 0.5–0.8 × 1–1.8 mm; horns usually convergent, 0.2–0.8 mm. |
turbinate to campanulate, 0.8–1 × 1.1–1.3 mm, glabrous or hairy; glands 4, green to yellow-green, oblong to subcircular, 0.2–0.4 × 0.4–0.6 mm; appendages absent or white, forming narrow rim at edge of gland, rarely slightly wider than gland, (0–)0.1(–0.3) × 0.4–0.6 mm, distal margin entire. |
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Staminate flowers | 8–12. |
8–14. |
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Pistillate flowers | ovary glabrous; styles 1–1.8 mm, 2-fid. |
ovary glabrous or hairy, subtended by triangular pad of tissue; styles spreading, 0.3–0.4 mm, 2-fid 1/2 to nearly entire length. |
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Capsules | depressed-ovoid, 4.3–5 × 5–6 mm, 3-lobed; cocci rounded, smooth, glabrous; columella 3.5–4 mm. |
broadly deltoid, 1.2–1.5 × 2–2.2 mm, glabrous or hairy; columella 0.9–1.3 mm. |
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Seeds | gray to dark brown, ovoid-oblong, truncate at both ends, 2.6–3.4 × 2–2.6 mm, shallowly pitted to almost smooth; caruncle conic, 1 × 0.8 mm. |
reddish brown, ovoid, 4-angled in cross section, 0.8–1.2 × 0.5–0.6 mm, obscurely wrinkled. |
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Cyathial | arrangement: terminal pleiochasial branches 3–5(–6), each 3–4 times 2-branched; pleiochasial bracts ovate-lanceolate to slightly subpandurate, similar in size or wider than distal leaves; dichasial bracts distinct, lanceolate to broadly ovate, base usually truncate to rounded or subcordate, sometimes attenuate, margins entire or slightly crenulate, apex usually obtuse to acute, occasionally acuminate; axillary cymose branches 2–8. |
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Cyathia | peduncle 1–3 mm. |
solitary at distal nodes; peduncle 0.7–1.5 mm. |
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2n | = 26. |
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Euphorbia chamaesula |
Euphorbia deltoidea |
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Phenology | Flowering and fruiting spring–summer. | |||||||||||||||||
Habitat | Clearings in ponderosa pine forests, montane roadsides, dry streambeds, creek banks, sandy and gravelly soils. | |||||||||||||||||
Elevation | 1700–2700 m. (5600–8900 ft.) | |||||||||||||||||
Distribution |
AZ; NM; Mexico (Chihuahua, Sonora)
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FL |
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Discussion | Euphorbia chamaesula is easily distinguished from other perennial members of subg. Esula in western North America by its larger capsules and the vegetative shoots (without cyathia) that arise from the distal nodes of the stem proximal to the pleiochasia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Subspecies 4 (4 in the flora). Euphorbia deltoidea comprises four narrowly endemic subspecies, all of which are endangered due to habitat loss and fragmentation. The subspecies occur in pine rockland habitat that is free of shrubby undergrowth. Periodic fires are required to keep the rockland habitat open. Subspecies serpyllum is restricted to Big Pine Key, Monroe County, whereas the other subspecies are found only in Miami-Dade County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 300. | FNA vol. 12, p. 265. | ||||||||||||||||
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum | ||||||||||||||||
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Synonyms | Tithymalus chamaesula | Chamaesyce deltoidea | ||||||||||||||||
Name authority | Boissier: Cent. Euphorb., 38. (1860) | Engelmann ex Chapman: Fl. South. U.S. ed. 2, 647. (1883) | ||||||||||||||||
Web links |