Euphorbia chamaesula |
Euphorbia astyla |
|
---|---|---|
mountain spurge |
alkali spurge, Pecos spurge |
|
Habit | Herbs, perennial, with thick rootstock. | Herbs, perennial, with woody or fibrous-fleshy taproot, 5–12 mm thick. |
Stems | erect, branched, 40–90 cm, glabrous. |
decumbent, ascending, or erect, few to many emerging from woody crown, 5–25(–50) cm, glabrous. |
Leaves | petiole 0.5–1 mm; blade elliptic to oblong, 8–20(–40) × 3–6 mm, base truncate, rounded, or attenuate, margins entire, apex obtuse or acute, sometimes slightly mucronate, surfaces glabrous; venation inconspicuous, only midvein prominent. |
opposite; stipules connate into deltate scale, 0.2–0.5 mm, minutely lacerate at apex, glabrous; petiole 0–0.2(–0.3) mm, glabrous; blade orbiculate-reniform to acute-cordate, 2–5(–8) × 2–5(–6) mm, base ± asymmetric, cordate to auriculate, sometimes clasping stem, margins entire, apex narrowly acute, surfaces glabrous; 2- or 3-veined from base, but usually only midvein conspicuous. |
Involucre | turbinate or campanulate, 1.5–2.5 × 1.1–1.9 mm, glabrous; glands 4, crescent-shaped to semicircular, 0.5–0.8 × 1–1.8 mm; horns usually convergent, 0.2–0.8 mm. |
broadly campanulate, 0.8–1.4 × 0.9–1.4 mm, glabrous; glands 4, yellow-green to brownish, oblong, 0.2–0.3 × 0.5–0.7 mm; appendages white, flabellate to oblong, 0.1–0.2(–0.5) × 0.4–0.8 mm, distal margin entire or dentate-crenate. |
Staminate flowers | 8–12. |
22–26. |
Pistillate flowers | ovary glabrous; styles 1–1.8 mm, 2-fid. |
ovary glabrous; styles 0.3–0.4 mm, unbranched, thickened-clavate. |
Capsules | depressed-ovoid, 4.3–5 × 5–6 mm, 3-lobed; cocci rounded, smooth, glabrous; columella 3.5–4 mm. |
ovoid and broadly triangular, 1.5–1.9(–2.5) × 1.4–1.6(–2.2) mm, glabrous; columella 1.2–1.8 mm. |
Seeds | gray to dark brown, ovoid-oblong, truncate at both ends, 2.6–3.4 × 2–2.6 mm, shallowly pitted to almost smooth; caruncle conic, 1 × 0.8 mm. |
white, oblong, 4-angled in cross section, adaxial faces slightly concave, with long raphe between, 1.5–1.8 × 0.7–1 mm, markedly foveolate, with irregular to ± parallel or anastomosing ridges. |
Cyathial | arrangement: terminal pleiochasial branches 3–5(–6), each 3–4 times 2-branched; pleiochasial bracts ovate-lanceolate to slightly subpandurate, similar in size or wider than distal leaves; dichasial bracts distinct, lanceolate to broadly ovate, base usually truncate to rounded or subcordate, sometimes attenuate, margins entire or slightly crenulate, apex usually obtuse to acute, occasionally acuminate; axillary cymose branches 2–8. |
|
Cyathia | peduncle 1–3 mm. |
solitary at distal nodes; peduncle 0.3–1(–1.5) mm. |
2n | = 26. |
= 28. |
Euphorbia chamaesula |
Euphorbia astyla |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting late spring–early fall. |
Habitat | Clearings in ponderosa pine forests, montane roadsides, dry streambeds, creek banks, sandy and gravelly soils. | Desert, grasslands, limestone substrates, usually on very saline or alkaline soils. |
Elevation | 1700–2700 m. (5600–8900 ft.) | 700–1100 m. (2300–3600 ft.) |
Distribution |
AZ; NM; Mexico (Chihuahua, Sonora)
|
TX; Mexico (Coahuila, Nuevo León) |
Discussion | Euphorbia chamaesula is easily distinguished from other perennial members of subg. Esula in western North America by its larger capsules and the vegetative shoots (without cyathia) that arise from the distal nodes of the stem proximal to the pleiochasia. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia astyla is a specialist on halophytic, alkaline soils and is known in the flora area only in part of Pecos County. The species is closely related to E. jejuna but differs in its sessile or sub-sessile leaves with a cordate-auriculate base and involucral gland appendages that are not deeply lobed or cleft. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 300. | FNA vol. 12, p. 261. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Tithymalus chamaesula | Chamaesyce astyla |
Name authority | Boissier: Cent. Euphorb., 38. (1860) | Engelmann ex Boissier: in A. P. de Candolle and A. L. P. P. de Candolle, Prodr. 15(2): 40. (1862) |
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