Euphorbia chaetocalyx |
Euphorbia prostrata |
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bristlecup sandmat |
blue weed, ground spurge, prostrate sandmat, prostrate spurge or sandmat |
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Habit | Herbs, perennial, with woody, thickened taproot. | Herbs, annual, with taproot. | ||||
Stems | usually erect, rarely slightly decumbent, often densely clustered from top of woody crown, 3–15 cm, glabrous. |
prostrate to decumbent, usually not mat-forming, 10–30 cm, crisped-villous to glabrate (proximally). |
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Leaves | opposite; stipules distinct, narrowly linear, usually entire, 0.5–1 mm, glabrous; petiole 0.5–1 mm, glabrous; blade ovate to lanceolate or oblong- or linear-lanceolate, 3–11 × 0.8–3(–5) mm, base slightly asymmetric, short-tapered, occasionally one side slightly rounded, margins entire, apex acute or short-acuminate, surfaces glabrous; only midvein conspicuous. |
opposite; stipules connate (lower side) or distinct (upper side), triangular-subulate, often lacerate distally, 0.5–1 mm, short crisped-villous to glabrate; petiole 0.5–1.5 mm, usually short crisped-villous, sometimes wooly at distal nodes; blade broadly elliptic to elliptic-oblong, ovate-spatulate, or ovate, 3–11(–15) × 3–6(–8) mm, base slightly asymmetric, rounded to slightly cordate and oblique, margins serrulate at least in distal 1/2, sometimes obscurely so, apex obtuse, abaxial surface finely crisped-villous, adaxial surface usually glabrous or glabrate, sometimes sparsely crisped-villous; 3-veined from base. |
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Involucre | campanulate to turbinate, 0.8–1.4 × 0.8–1 mm, glabrous; glands 4, yellow-brown to reddish, concave or convex, elliptic or oval, 0.2–0.4 × 0.4–0.6 mm; appendages absent or white, lanceolate-deltate to straplike, 0.2–1.1 × 0.2–0.9 mm, distal margin entire, crenate, or deeply cleft or divided. |
obconic, 0.6–0.9 × 0.5 mm, crisped-villous or glabrous; glands 4, reddish, oval to oblong, 0.1 × 0.1–0.2 mm; appendages white to pink, rudimentary, 0–0.2 mm, distal margin entire or irregularly scalloped. |
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Staminate flowers | 25–35. |
3–6. |
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Pistillate flowers | ovary glabrous; styles 0.3–0.4 mm, 2-fid 1/2 length. |
ovary densely crisped-villous; styles 0.1 mm, 2-fid nearly entire length. |
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Capsules | depressed-ovoid to depressed-globose, 1.7–2.1 × 1.6–2.4 mm, glabrous; columella 1.2–1.8 mm. |
broadly ovoid, 1.2–2 × 1.4–1.5 mm, crisped-villous along keels and toward base, often glabrous between keels; columella 1–1.2 mm. |
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Seeds | white, ovoid-pyramidal, prominently 4-angled in cross section, 1.6–2 × 1–1.2 mm, smooth to slightly wrinkled. |
white but with barely concealed brown surface beneath, ovoid, sharply 4-angled in cross section, abaxial faces plane to convex, adaxial faces concave, 0.8–1.1 × 0.5–0.7 mm, with several narrow, sharp, slightly irregular, transverse ridges. |
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Cyathia | solitary at distal nodes; peduncle 0.8–1.3 mm. |
solitary or in small, cymose clusters at distal nodes or on congested, axillary branches; peduncle 1–2 mm. |
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2n | = 18. |
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Euphorbia chaetocalyx |
Euphorbia prostrata |
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Phenology | Flowering and fruiting early spring–fall. | |||||
Habitat | Disturbed areas, fields, gardens, sidewalks, sandy places, ballast piles. | |||||
Elevation | 0–1400 m. (0–4600 ft.) | |||||
Distribution |
AZ; NM; TX; n Mexico
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AL; AR; AZ; CA; CO; FL; GA; IA; IL; IN; KS; KY; LA; MA; MD; MI; MO; MS; NC; NE; NM; OH; OK; PA; SC; SD; TN; TX; UT; VA; WV; WY; Mexico; Central America; South America; West Indies [Introduced in Eurasia, Africa, Pacific Islands, Australia]
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Discussion | Varieties 2 (2 in the flora). Euphorbia chaetocalyx is similar to E. fendleri but can generally be distinguished from that species by its narrow, acute leaves and ± erect stems. Some authors have used the presence or absence and shape of the involucral gland appendages to help separate E. chaetocalyx from E. fendleri, but those characters appear highly variable and of little taxonomic utility. Some individuals from western Texas (Culberson and El Paso counties) and southern New Mexico appear intermediate with E. fendleri. The specific epithet of E. chaetocalyx refers to the bristly perianthlike segments that subtend the ovary, but these structures are found intermittently in both E. chaetocalyx and E. fendleri. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia prostrata is native to tropical America and possibly into the southern part of the flora area. It is likely adventive throughout most of the northern part of its range. It is widely naturalized throughout much of the rest of the tropics. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 263. | FNA vol. 12, p. 285. | ||||
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | E. fendleri var. chaetocalyx, Chamaesyce chaetocalyx | Chamaesyce prostrata | ||||
Name authority | (Boissier) Tidestrom: Proc. Biol. Soc. Wash. 48: 40. (1935) | Aiton: Hort. Kew. 2: 139. (1789) | ||||
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