Euphorbia chaetocalyx |
Euphorbia glyptosperma |
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bristlecup sandmat |
corrugate-seed spurge, rib-seed sandmat, ridge-seed spurge |
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Habit | Herbs, perennial, with woody, thickened taproot. | Herbs, annual, with taproot. | ||||
Stems | usually erect, rarely slightly decumbent, often densely clustered from top of woody crown, 3–15 cm, glabrous. |
prostrate, loosely mat-forming, 5–40 cm, glabrous, sometimes slightly glaucous. |
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Leaves | opposite; stipules distinct, narrowly linear, usually entire, 0.5–1 mm, glabrous; petiole 0.5–1 mm, glabrous; blade ovate to lanceolate or oblong- or linear-lanceolate, 3–11 × 0.8–3(–5) mm, base slightly asymmetric, short-tapered, occasionally one side slightly rounded, margins entire, apex acute or short-acuminate, surfaces glabrous; only midvein conspicuous. |
opposite; stipules usually distinct, linear-subulate, usually irregularly fringed or lobed, rarely laciniate, 0.4–2 mm, glabrous; petiole 0.2–2 mm, glabrous; blade narrowly oblong to oblong-obovate or oblong-ovate, 3–15 × 2–7 mm, base asymmetric, one side angled and other rounded to shallowly cordate, margins minutely sparsely serrulate, especially near apex and on longer side, apex rounded to obtuse, abaxial surface usually pale grayish green, adaxial surface sometimes reddish tinged or with reddish blotch, both surfaces glabrous; palmately veined at base, pinnate distally. |
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Involucre | campanulate to turbinate, 0.8–1.4 × 0.8–1 mm, glabrous; glands 4, yellow-brown to reddish, concave or convex, elliptic or oval, 0.2–0.4 × 0.4–0.6 mm; appendages absent or white, lanceolate-deltate to straplike, 0.2–1.1 × 0.2–0.9 mm, distal margin entire, crenate, or deeply cleft or divided. |
obconic, 0.6–0.9 × 0.3–0.6 mm, glabrous; glands 4, red to purple, narrowly oblong-elliptic, 0.1–0.2 × 0.1–0.5 mm; appendages white or pinkish tinged, semilunate and fringing distal margin of gland, 0.1–0.3 × 0.1–0.3 mm, distal margin usually irregularly crenulate or lobed. |
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Staminate flowers | 25–35. |
1–5. |
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Pistillate flowers | ovary glabrous; styles 0.3–0.4 mm, 2-fid 1/2 length. |
ovary glabrous; styles 0.1–0.3 mm, 2-fid 1/2 length. |
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Capsules | depressed-ovoid to depressed-globose, 1.7–2.1 × 1.6–2.4 mm, glabrous; columella 1.2–1.8 mm. |
broadly ovoid, 1.3–1.9 × 1.6–2 mm, glabrous; columella 1.3–1.5 mm. |
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Seeds | white, ovoid-pyramidal, prominently 4-angled in cross section, 1.6–2 × 1–1.2 mm, smooth to slightly wrinkled. |
with thin white coat that readily falls off, surface below tan brown, oblong-ovoid, sharply angular in cross section, 1–1.4 × 0.6–0.9 mm, with 3–4(–6) prominent transverse ridges that usually interrupt abaxial keel. |
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Cyathia | solitary at distal nodes; peduncle 0.8–1.3 mm. |
solitary or in small, cymose clusters at distal nodes; peduncle 0.5–1.6 mm. |
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2n | = 22. |
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Euphorbia chaetocalyx |
Euphorbia glyptosperma |
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Phenology | Flowering and fruiting early summer–fall. | |||||
Habitat | Stream and river banks, sand prairies, loess hill prairies, meadows, ballast, open disturbed areas, roadsides. | |||||
Elevation | 0–1800 m. (0–5900 ft.) | |||||
Distribution |
AZ; NM; TX; n Mexico
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AZ; CA; CO; CT; IA; ID; IL; IN; KS; LA; MA; ME; MI; MN; MO; MT; ND; NE; NH; NM; NV; NY; OK; OR; SD; TN; TX; UT; VT; WA; WI; WY; AB; BC; MB; NB; NS; ON; PE; QC; SK; Mexico (Chihuahua, Coahuila, Tamaulipas, Zacatecas)
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Discussion | Varieties 2 (2 in the flora). Euphorbia chaetocalyx is similar to E. fendleri but can generally be distinguished from that species by its narrow, acute leaves and ± erect stems. Some authors have used the presence or absence and shape of the involucral gland appendages to help separate E. chaetocalyx from E. fendleri, but those characters appear highly variable and of little taxonomic utility. Some individuals from western Texas (Culberson and El Paso counties) and southern New Mexico appear intermediate with E. fendleri. The specific epithet of E. chaetocalyx refers to the bristly perianthlike segments that subtend the ovary, but these structures are found intermittently in both E. chaetocalyx and E. fendleri. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia glyptosperma is one of the most widespread species of the genus in North America, and it is often quite weedy. It is likely native to much of the central United States, but in areas like eastern Canada, it occurs in highly disturbed habitats, and it may be adventive there. Euphorbia glyptosperma is characterized by its prominently ridged seeds, laciniate stipules, complete lack of pubescence, and somewhat obscure toothing near the tips of the leaves. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 263. | FNA vol. 12, p. 269. | ||||
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | E. fendleri var. chaetocalyx, Chamaesyce chaetocalyx | Chamaesyce glyptosperma | ||||
Name authority | (Boissier) Tidestrom: Proc. Biol. Soc. Wash. 48: 40. (1935) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 187. (1859) | ||||
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