Euphorbia chaetocalyx |
Euphorbia garberi |
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bristlecup sandmat |
Garber's sandmat |
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Habit | Herbs, perennial, with woody, thickened taproot. | Herbs, perennial, with moderately thickened rootstock. | ||||
Stems | usually erect, rarely slightly decumbent, often densely clustered from top of woody crown, 3–15 cm, glabrous. |
ascending, sometimes slightly woody at base, not wiry, 1–3 mm diam., 15–50 cm, canescent. |
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Leaves | opposite; stipules distinct, narrowly linear, usually entire, 0.5–1 mm, glabrous; petiole 0.5–1 mm, glabrous; blade ovate to lanceolate or oblong- or linear-lanceolate, 3–11 × 0.8–3(–5) mm, base slightly asymmetric, short-tapered, occasionally one side slightly rounded, margins entire, apex acute or short-acuminate, surfaces glabrous; only midvein conspicuous. |
opposite; stipules distinct or connate at base, triangular-subulate, apex divided into 2–5 subulate to subulate-filiform segments, 0.5–0.7 mm, pilose; petiole 0.5–1.1 mm, pilose; blade ovate to oblong-elliptic, 4–9(–15) × 3–6 mm, base asymmetric, obtuse to rounded, margins usually entire, rarely very sparsely serrulate, apex usually obtuse to rounded, occasionally acute to apiculate, surfaces canescent; 3-veined from base, only midvein conspicuous. |
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Involucre | campanulate to turbinate, 0.8–1.4 × 0.8–1 mm, glabrous; glands 4, yellow-brown to reddish, concave or convex, elliptic or oval, 0.2–0.4 × 0.4–0.6 mm; appendages absent or white, lanceolate-deltate to straplike, 0.2–1.1 × 0.2–0.9 mm, distal margin entire, crenate, or deeply cleft or divided. |
turbinate to campanulate, 0.6–0.9 × 0.5–0.9 mm, pilose; glands 4, brown, slightly concave, elliptic to subcircular, 0.1–0.3 × 0.2–0.4 mm; appendages white to pink, as narrow rim along gland, (0–)0.1–0.2 × 0.3–0.5 mm, surfaces pilose, distal margin entire or crenulate. |
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Staminate flowers | 25–35. |
8–20. |
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Pistillate flowers | ovary glabrous; styles 0.3–0.4 mm, 2-fid 1/2 length. |
ovary pilose; styles 0.6–0.7 mm, 2-fid 1/2 length. |
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Capsules | depressed-ovoid to depressed-globose, 1.7–2.1 × 1.6–2.4 mm, glabrous; columella 1.2–1.8 mm. |
subglobose to broadly ovoid, 1.1–1.6 × 1.3–2.1 mm, pilose; columella 1.1–1.6 mm. |
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Seeds | white, ovoid-pyramidal, prominently 4-angled in cross section, 1.6–2 × 1–1.2 mm, smooth to slightly wrinkled. |
gray to reddish brown, oblong-ovoid, 4-angled in cross section, 1–1.2 × 0.6–0.8 mm, flat or obscurely wrinkled. |
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Cyathia | solitary at distal nodes; peduncle 0.8–1.3 mm. |
solitary at nodes of short, axillary branches; peduncle 0.2–0.4 mm. |
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Euphorbia chaetocalyx |
Euphorbia garberi |
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Phenology | Flowering and fruiting year-round. | |||||
Habitat | Beach dunes, coastal rock barrens, disturbed uplands and pine rocklands. | |||||
Elevation | 0–10 m. (0–0 ft.) | |||||
Distribution |
AZ; NM; TX; n Mexico
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FL |
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Discussion | Varieties 2 (2 in the flora). Euphorbia chaetocalyx is similar to E. fendleri but can generally be distinguished from that species by its narrow, acute leaves and ± erect stems. Some authors have used the presence or absence and shape of the involucral gland appendages to help separate E. chaetocalyx from E. fendleri, but those characters appear highly variable and of little taxonomic utility. Some individuals from western Texas (Culberson and El Paso counties) and southern New Mexico appear intermediate with E. fendleri. The specific epithet of E. chaetocalyx refers to the bristly perianthlike segments that subtend the ovary, but these structures are found intermittently in both E. chaetocalyx and E. fendleri. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia garberi appears to be an interspecific hybrid, but its parentage is not entirely clear (Y. Yang and P. E. Berry 2011). The taxon is closely related to E. blodgettii, E. porteriana, and E. serpens. It is restricted to Collier, Miami-Dade, and Monroe counties in southern Florida and is federally listed as threatened due to impacts from habitat loss, fire suppression, and invasive species. A. Herndon (1993) synonymized E. porteriana var. keyensis under E. garberi because of its uniformly pilose capsules, and that treatment is followed here. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Key |
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Source | FNA vol. 12, p. 263. | FNA vol. 12, p. 268. | ||||
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum | ||||
Sibling taxa | ||||||
Subordinate taxa | ||||||
Synonyms | E. fendleri var. chaetocalyx, Chamaesyce chaetocalyx | Chamaesyce adicioides, C. brachypoda, C. garberi, C. keyensis, C. mosieri, C. porteriana var. keyensis, E. porteriana var. keyensis | ||||
Name authority | (Boissier) Tidestrom: Proc. Biol. Soc. Wash. 48: 40. (1935) | Engelmann ex Chapman: Fl. South. U.S. ed. 2, 646. (1883) | ||||
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