Euphorbia capitellata |
Euphorbia tetrapora |
|
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capitate sandmat, head sandmat, head spurge |
weak spurge |
|
Habit | Herbs, annual or perennial, with slender to thick and woody rootstock. | Herbs, annual, with taproot. |
Stems | usually ascending (but ranging from decumbent to erect), 15–50 cm, glabrous, strigillose, or pilose. |
erect, unbranched, 7–20 cm, glabrous. |
Leaves | opposite; stipules distinct, filiform or divided into 2–3 subulate-filiform segments, without dark, circular glands at base, 0.6–1.5 mm, pilose; petioles 0.6–1.3 mm, glabrous, pilose, or strigillose; blade ovate to narrowly ovate, 4–19 × 2–8 mm, base asymmetric, one side strongly cordate, other side rounded to slightly cordate, margins entire or serrulate (commonly nearly entire with few scattered teeth, often slightly thickened), apex acute, surfaces often with red spot in center, glabrous, pilose, or strigillose; weakly 3-veined from base, usually only midvein conspicuous. |
petiole 1–2 mm, reduced distally; blade spatulate-cuneate, 8–10 × 4–5 mm (greatly reduced in size proximally), base cuneate, margins entire, apex rounded to emarginate or obcordate, surfaces glabrous; venation pinnate. |
Involucre | narrowly obconic to narrowly campanulate, 0.8–1.6 × 0.7–1.3 mm, glabrous or pilose; glands 4, yellow-green to pink or maroon, circular to oblong, 0.2–0.4 × 0.2–0.5 mm; appendages white to light pink, oblong to reniform or flabellate, 0.2–1.1 × 0.5–1.7 mm, surfaces glabrous, distal margin entire. |
campanulate, 0.8–1.1 × 0.7–1 mm, glabrous; glands 4, elliptic to trapezoidal, 0.3–0.6 × 0.6–1.2 mm; horns divergent, 0.5–1 mm. |
Staminate flowers | 25–40. |
10–15. |
Pistillate flowers | ovary glabrous or pilose; styles 0.4–0.6 mm, 2-fid entire length. |
ovary smooth, glabrous; styles 0.6–1 mm, 2-fid. |
Capsules | ovoid to oblate, 1.3–1.9 × 1.4–2.1 mm, glabrous or pilose; columella 1.1–1.7 mm. |
depressed-globose, 1.8–2.2 × 2.2–2.9 mm, slightly lobed; cocci rounded to slightly flattened, smooth, glabrous; columella 1.5–1.8 mm. |
Seeds | pink to pinkish gray, narrowly ovoid to narrowly ovoid-oblong, 4-angled or weakly 3-angled in cross section, 0.9–1.5 × 0.5–0.7 mm, irregularly dimpled, sometimes also with faint transverse ridges that do not pass through abaxial keel. |
reddish brown to brown, often glaucous, oblong, 1.3–1.4 × 0.8–0.9 mm, abaxial faces with 15–20 shallow pits or almost smooth, adaxial faces with 4–6 large shallow pits or irregular oblong grooves; caruncle conic, hat-shaped, 0.3–0.4 × 0.4–0.6 mm. |
Cyathia | in dense, terminal capitate glomerules, with reduced, bractlike leaves subtending cyathia, at tips of main stems and short, leafy, axillary branches; peduncle 0.1–1.2 mm. |
peduncle 0.2–0.6 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3, each 1–3(–4) times 2-branched; pleiochasial bracts obovate, similar in size to distal leaves; dichasial bracts distinct or basally subconnate, not imbricate, triangular-ovate, base truncate or cordate, margins entire, apex mucronate; axillary cymose branches 1–4. |
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2n | = 14. |
|
Euphorbia capitellata |
Euphorbia tetrapora |
|
Phenology | Flowering and fruiting year-round in response to sufficient moisture. | Flowering and fruiting spring. |
Habitat | Gravelly washes, rocky slopes, basaltic talus, disturbed roadsides, primarily desert scrub, desert grasslands, riparian forests, rarely oak-juniper woodlands. | Sandy soils, dry open woods. |
Elevation | 600–1600 m. (2000–5200 ft.) | 0–300 m. (0–1000 ft.) |
Distribution |
AZ; NM; TX; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Sinaloa, Sonora)
|
LA; OK; TX |
Discussion | Euphorbia capitellata is a characteristic herb in the Sonoran Desert of southern Arizona, ranging east to extreme southwestern Texas. During peak flowering, plants are attractive due to the dense clusters of cyathia with well-developed involucral gland appendages. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia tetrapora is endemic to a portion of the western Gulf coastal plain. D. S. Correll and M. C. Johnston (1970) included Alabama and Georgia in the distribution of this species as well, probably due to Engelmann’s citation of a Georgia specimen from the herbarium of Samuel Boykin. Whether the Boykin specimen came from Georgia, where Boykin was based, is unclear. Because no records to support its occurrence in the eastern Gulf coastal plain (Alabama or Georgia) have been found, those states are here excluded from the distribution of E. tetrapora. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 262. | FNA vol. 12, p. 311. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | Chamaesyce capitellata, C. pycnanthema, E. pycnanthema | Tithymalus tetraporus |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 188. (1859) | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 191. (1859) |
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