Euphorbia capitellata |
Euphorbia mendezii |
|
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capitate sandmat, head sandmat, head spurge |
Mendez's sandmat |
|
Habit | Herbs, annual or perennial, with slender to thick and woody rootstock. | Herbs, annual or perennial, with slender taproot. |
Stems | usually ascending (but ranging from decumbent to erect), 15–50 cm, glabrous, strigillose, or pilose. |
prostrate, often mat-forming, 8–35 cm, usually villous along margins, lower surface glabrous, upper surface usually strigillose to puberulent, rarely glabrous or glabrate. |
Leaves | opposite; stipules distinct, filiform or divided into 2–3 subulate-filiform segments, without dark, circular glands at base, 0.6–1.5 mm, pilose; petioles 0.6–1.3 mm, glabrous, pilose, or strigillose; blade ovate to narrowly ovate, 4–19 × 2–8 mm, base asymmetric, one side strongly cordate, other side rounded to slightly cordate, margins entire or serrulate (commonly nearly entire with few scattered teeth, often slightly thickened), apex acute, surfaces often with red spot in center, glabrous, pilose, or strigillose; weakly 3-veined from base, usually only midvein conspicuous. |
opposite; stipules usually distinct or connate basally, rarely completely connate, deltate, laciniate, glabrous or pilose (lower side), forming narrow deltate scale, sometimes apically 2-fid or laciniate, glabrous (upper side), 0.4–1.9 mm; petiole 0.3–1.2 mm, glabrous, pilose or villous; blade oblong to obovate, 4–12 × 2–7 mm, base asymmetric, one side attenuate, cuneate or rounded, other rounded or cordate, margins serrulate at least distally, apex obtuse, surfaces glabrous or sparsely sericeous, pilose or villous; 3-veined at base. |
Involucre | narrowly obconic to narrowly campanulate, 0.8–1.6 × 0.7–1.3 mm, glabrous or pilose; glands 4, yellow-green to pink or maroon, circular to oblong, 0.2–0.4 × 0.2–0.5 mm; appendages white to light pink, oblong to reniform or flabellate, 0.2–1.1 × 0.5–1.7 mm, surfaces glabrous, distal margin entire. |
campanulate or obconic, 0.8–1 × 0.7–0.8 mm, usually glabrous, rarely sparsely pilose toward apex; glands 4, pink, reniform, oblong or elliptic, 0.1 × 0.2–0.3 mm; appendages absent or white to pink, oblong, flabellate or forming narrow rim around edge of gland, 0.1–0.3 × 0.3–0.6 mm, distal margin usually entire, sometimes lobed. |
Staminate flowers | 25–40. |
6–15. |
Pistillate flowers | ovary glabrous or pilose; styles 0.4–0.6 mm, 2-fid entire length. |
ovary usually pilose or villous with hairs concentrated along keels, rarely glabrous; styles 0.2–0.3 mm, 2-fid 1/2 length. |
Capsules | ovoid to oblate, 1.3–1.9 × 1.4–2.1 mm, glabrous or pilose; columella 1.1–1.7 mm. |
ovoid, 1.2–1.6 × 1.2–1.4 mm, usually pilose or villous with hairs concentrated along keels, often glabrous in between, very rarely completely glabrous; columella 1–1.4 mm. |
Seeds | pink to pinkish gray, narrowly ovoid to narrowly ovoid-oblong, 4-angled or weakly 3-angled in cross section, 0.9–1.5 × 0.5–0.7 mm, irregularly dimpled, sometimes also with faint transverse ridges that do not pass through abaxial keel. |
reddish brown to orange or gray-pink, narrowly ovoid, 4-angled in cross section, 0.9–1.2 × 0.5–0.6 mm, almost smooth or with 5–7 faint transverse ridges that do not pass through abaxial keel. |
Cyathia | in dense, terminal capitate glomerules, with reduced, bractlike leaves subtending cyathia, at tips of main stems and short, leafy, axillary branches; peduncle 0.1–1.2 mm. |
solitary at nodes or on short, congested axillary branches; peduncle 0.9–2.5 mm. |
2n | = 14. |
|
Euphorbia capitellata |
Euphorbia mendezii |
|
Phenology | Flowering and fruiting year-round in response to sufficient moisture. | Flowering and fruiting year-round. |
Habitat | Gravelly washes, rocky slopes, basaltic talus, disturbed roadsides, primarily desert scrub, desert grasslands, riparian forests, rarely oak-juniper woodlands. | Disturbed areas. |
Elevation | 600–1600 m. (2000–5200 ft.) | 0–10 m. (0–0 ft.) |
Distribution |
AZ; NM; TX; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Sinaloa, Sonora)
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FL; Mexico; Central America; West Indies; n South America [Introduced in North America] |
Discussion | Euphorbia capitellata is a characteristic herb in the Sonoran Desert of southern Arizona, ranging east to extreme southwestern Texas. During peak flowering, plants are attractive due to the dense clusters of cyathia with well-developed involucral gland appendages. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia mendezii is a common weed distributed widely throughout Mexico and Central America. Within the flora area the species is known only from southern Florida. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 262. | FNA vol. 12, p. 277. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce capitellata, C. pycnanthema, E. pycnanthema | Chamaesyce mendezii |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 188. (1859) | Boissier: Cent. Euphorb., 15. (1860) |
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