Euphorbia capitellata |
Euphorbia abramsiana |
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capitate sandmat, head sandmat, head spurge |
Abrams' sandmat, Abrams' spurge |
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Habit | Herbs, annual or perennial, with slender to thick and woody rootstock. | Herbs, annual, with slender taproot. |
Stems | usually ascending (but ranging from decumbent to erect), 15–50 cm, glabrous, strigillose, or pilose. |
prostrate, mat-forming, 10–35(–50) cm, shortly pilose or puberulent at least proximally, often glabrous distally. |
Leaves | opposite; stipules distinct, filiform or divided into 2–3 subulate-filiform segments, without dark, circular glands at base, 0.6–1.5 mm, pilose; petioles 0.6–1.3 mm, glabrous, pilose, or strigillose; blade ovate to narrowly ovate, 4–19 × 2–8 mm, base asymmetric, one side strongly cordate, other side rounded to slightly cordate, margins entire or serrulate (commonly nearly entire with few scattered teeth, often slightly thickened), apex acute, surfaces often with red spot in center, glabrous, pilose, or strigillose; weakly 3-veined from base, usually only midvein conspicuous. |
opposite; stipules distinct, divided into 5–7 subulate-filiform segments, 0.6–1.1 mm, usually glabrous, rarely pilose; petiole 0.5–1 mm, glabrous; blade ovate, elliptic-oblong, or slightly ovate-cordate, 3–11 × 2–5 mm, base asymmetric, truncate to hemicordate, margins serrulate at least toward apex, often entire toward base, apex acute to obtuse, surfaces sometimes with red spot in center, glabrous; usually only the midvein conspicuous. |
Involucre | narrowly obconic to narrowly campanulate, 0.8–1.6 × 0.7–1.3 mm, glabrous or pilose; glands 4, yellow-green to pink or maroon, circular to oblong, 0.2–0.4 × 0.2–0.5 mm; appendages white to light pink, oblong to reniform or flabellate, 0.2–1.1 × 0.5–1.7 mm, surfaces glabrous, distal margin entire. |
obconic, 0.5–0.6 × 0.4–0.5 mm, glabrous; glands 4, yellowish to pink, circular to oblong, 0.1 × 0.1–0.2 mm; appendages absent, or white to pink, semicircular to broadly ovate, to 0.1 × 0.2 mm, distal margin entire or shallowly lobed. |
Staminate flowers | 25–40. |
3–5. |
Pistillate flowers | ovary glabrous or pilose; styles 0.4–0.6 mm, 2-fid entire length. |
ovary glabrous; styles 0.1–0.3 mm, 2-fid nearly entire length. |
Capsules | ovoid to oblate, 1.3–1.9 × 1.4–2.1 mm, glabrous or pilose; columella 1.1–1.7 mm. |
ellipsoid to ovoid, 1.3–1.5 × 1.1–1.5 mm, glabrous; columella 1–1.3 mm. |
Seeds | pink to pinkish gray, narrowly ovoid to narrowly ovoid-oblong, 4-angled or weakly 3-angled in cross section, 0.9–1.5 × 0.5–0.7 mm, irregularly dimpled, sometimes also with faint transverse ridges that do not pass through abaxial keel. |
light gray to light brown, narrowly ovoid to ovoid, 4-angled in cross section, 1–1.2 × 0.6–0.7 mm, with 3–5 prominent transverse ridges that often interrupt abaxial keel. |
Cyathia | in dense, terminal capitate glomerules, with reduced, bractlike leaves subtending cyathia, at tips of main stems and short, leafy, axillary branches; peduncle 0.1–1.2 mm. |
solitary at distal nodes of primary stems or at nodes of short congested axillary branchlets; peduncle 0.2–0.5 mm. |
2n | = 14. |
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Euphorbia capitellata |
Euphorbia abramsiana |
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Phenology | Flowering and fruiting year-round in response to sufficient moisture. | Flowering and fruiting summer–fall. |
Habitat | Gravelly washes, rocky slopes, basaltic talus, disturbed roadsides, primarily desert scrub, desert grasslands, riparian forests, rarely oak-juniper woodlands. | Desert scrub and desert grasslands. |
Elevation | 600–1600 m. (2000–5200 ft.) | -40–1400 m. (-100–4600 ft.) |
Distribution |
AZ; NM; TX; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Sinaloa, Sonora)
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AZ; CA; NM; TX; Mexico (Baja California, Baja California Sur, Sonora)
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Discussion | Euphorbia capitellata is a characteristic herb in the Sonoran Desert of southern Arizona, ranging east to extreme southwestern Texas. During peak flowering, plants are attractive due to the dense clusters of cyathia with well-developed involucral gland appendages. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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Source | FNA vol. 12, p. 262. | FNA vol. 12, p. 259. |
Parent taxa | Euphorbiaceae > Euphorbia > sect. Anisophyllum | Euphorbiaceae > Euphorbia > sect. Anisophyllum |
Sibling taxa | ||
Synonyms | Chamaesyce capitellata, C. pycnanthema, E. pycnanthema | Chamaesyce abramsiana |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 188. (1859) | L. C. Wheeler: Bull. S. Calif. Acad. Sci. 33: 109. (1934) |
Web links |