Euphorbia brachycera |
Euphorbia trichotoma |
|
---|---|---|
horned spurge, shorthorn spurge |
sand-dune spurge |
|
Habit | Herbs, perennial, with thick rootstock. | Herbs, perennial, with thick rootstock. |
Stems | erect or ascending, branched, 10–60 cm, usually glabrous, sometimes puberulent. |
erect, branched, 15–40 cm, glabrous. |
Leaves | petiole 0–0.5 mm; blade oblong-elliptic, lanceolate, or oblanceolate to broadly ovate, 5–25 × 2–7 mm, base truncate, rounded, acute, or attenuate, margins entire, apex acute or obtuse, sometimes mucronate, surfaces usually glabrous, sometimes puberulent; venation inconspicuous, only midvein prominent. |
petiole absent; blade obovate to oblanceolate, 5–20 × 2–7 mm, base broadly attenuate, margins entire, apex acute to obtuse, surfaces glabrous; venation pinnate, very obscure, midvein prominent. |
Involucre | turbinate, campanulate or cupulate, 1.5–2.6 × 1.4–1.7 mm, usually glabrous, sometimes puberulent; glands 4, crescent-shaped to triangular-ovate, 0.5–0.8 × 0.7–1.7 mm, margins usually entire, occasionally slightly crenate to dentate; horns divergent, 0.2–0.4 mm, longer than teeth on gland margin. |
hemispheric, 1–1.5 × 1.4–2 mm, glabrous; glands 5, elliptic to slightly crescent-shaped, 0.4–0.6 × 0.9–1.1 mm; horns divergent, 0.1–0.2 mm. |
Staminate flowers | 10–15. |
8–10. |
Pistillate flower(s) | ovary, usually glabrous, sometimes puberulent; styles 0.5–0.9 mm, 2-fid. |
ovary glabrous; styles 0.2–0.4 mm, 2-fid. |
Capsules | depressed-ovoid, 2.8–4 × 3.5–4.5 mm, 3-lobed; cocci rounded, smooth, usually glabrous, sometimes sparsely puberulent; columella 2.4–3.3 mm. |
depressed-ovoid, 1.1–1.5 × 2–3 mm, strongly 3-lobed; cocci rounded, minutely papillate, sometimes appearing smooth, glabrous; columella 1.6–1.9 mm. |
Seeds | light gray, cylindric ovoid-oblong, 2–2.8(–3) × 1.4–2.2 mm, irregularly shallowly pitted; caruncle sessile to shortly stipitate, conic, 0.4–0.5 × 0.8–1.1 mm. |
white, ovoid-globose, 1.8–2 × 1.6–1.8 mm, smooth; caruncle ± rounded and flattened, 0.3 × 0.3 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3–5(–8), 1–4+ times 2-branched; pleiochasial bracts lanceolate or ovate to broadly ovate, wider than distal leaves; dichasial bracts distinct, broadly ovate or rhombic to triangular-ovate, base obtuse, margins entire or slightly crenate, apex obtuse, mucronate; axillary cymose branches 0–8. |
arrangement: terminal pleiochasial branches 3, each usually 1–2 times 3-branched, sometimes unbranched; pleiochasial bracts similar in shape and size to distal leaves, alternate; dichasial bracts distinct, obovate to oblanceolate, base broadly attenuate, margins entire, apex acute to obtuse; axillary cymose branches 0. |
Cyathia | peduncle 0.4–1 mm. |
peduncle 0.7–1 mm. |
2n | = 28. |
|
Euphorbia brachycera |
Euphorbia trichotoma |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting year-round (primarily spring–summer). |
Habitat | Montane areas, canyons, rock crevices, sandy or gravelly slopes, pine-oak woodlands, ponderosa pine and mixed coniferous forests. | Coastal beaches, sand dunes, thickets. |
Elevation | 1200–3200 m. (3900–10500 ft.) | 0–10 m. (0–0 ft.) |
Distribution |
AZ; CO; ID; MT; ND; NE; NM; SD; TX; UT; WY; Mexico (Chihuahua, Coahuila, Sonora)
|
FL; Mexico; Central America; West Indies |
Discussion | Euphorbia brachycera is morphologically extremely variable, and plants from the northern part of the species range have sometimes been distinguished as E. robusta. Euphorbia brachycera is most closely related to the other native, perennial species of subg. Esula in western North America, namely E. chamaesula, E. lurida, E. schizoloba, and E. yaquiana. The name E. montana Engelmann, which has sometimes been applied to E. brachycera, is illegitimate because it is a later homonym of E. montana Rafinesque. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia trichotoma is found in coastal peninsular Florida from Hillsborough County south to Key West. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 300. | FNA vol. 12, p. 311. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | E. brachycera var. robusta, E. montana var. robusta, E. odontadenia, E. robusta, Tithymalus brachycerus, T. robustus | Galarhoeus trichotomus, Tithymalus trichotomus |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) | Kunth: in A. von Humboldt et al., Nov. Gen. Sp. 2(fol.): 48; 2(qto.): 60. (1817) |
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