Euphorbia brachycera |
Euphorbia telephioides |
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horned spurge, shorthorn spurge |
telephus spurge |
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Habit | Herbs, perennial, with thick rootstock. | Herbs, perennial, with thickened rootstock. |
Stems | erect or ascending, branched, 10–60 cm, usually glabrous, sometimes puberulent. |
erect or ascending, 20–30 cm. |
Leaves | petiole 0–0.5 mm; blade oblong-elliptic, lanceolate, or oblanceolate to broadly ovate, 5–25 × 2–7 mm, base truncate, rounded, acute, or attenuate, margins entire, apex acute or obtuse, sometimes mucronate, surfaces usually glabrous, sometimes puberulent; venation inconspicuous, only midvein prominent. |
petiole usually absent or indistinct, occasionally to 5.5 mm, blade elliptic or obovate, 31–60 × 7–32 mm, thick and fleshy, base attenuate or cuneate, apex acute, obtuse, or mucronate; venation pinnate with 8–13 lateral veins, these sometimes obscure and only midvein evident. |
Involucre | turbinate, campanulate or cupulate, 1.5–2.6 × 1.4–1.7 mm, usually glabrous, sometimes puberulent; glands 4, crescent-shaped to triangular-ovate, 0.5–0.8 × 0.7–1.7 mm, margins usually entire, occasionally slightly crenate to dentate; horns divergent, 0.2–0.4 mm, longer than teeth on gland margin. |
campanulate, 1.3–2.3 × 1.6–2.5 mm, lobes ovate to oblong, 0.5–0.7 mm, ciliate; glands purple-red, oblong to trapezoidal, 0.5–0.7 × 0.8–1.2 mm, distal margins entire or crenulate. |
Staminate flowers | 10–15. |
25–30. |
Pistillate flower(s) | ovary, usually glabrous, sometimes puberulent; styles 0.5–0.9 mm, 2-fid. |
gynophore exserted 1.9–3.2 mm, calyxlike lobes triangular, 0.2–0.7 mm; styles connate 1/4 length, 1.3–1.5 mm. |
Capsules | depressed-ovoid, 2.8–4 × 3.5–4.5 mm, 3-lobed; cocci rounded, smooth, usually glabrous, sometimes sparsely puberulent; columella 2.4–3.3 mm. |
ovoid-oblate, 5.2–5.6 × 6.6–8.3 mm, 3-lobed; columella 3.9–4.1 mm. |
Seeds | light gray, cylindric ovoid-oblong, 2–2.8(–3) × 1.4–2.2 mm, irregularly shallowly pitted; caruncle sessile to shortly stipitate, conic, 0.4–0.5 × 0.8–1.1 mm. |
blackish to dark brown, ovoid to globose-ovoid, circular in cross section, 3.2–3.5 × 2.6–3.1 mm, smooth, base rounded to flattened, apex rounded, occasionally with low point. |
Cyathial | arrangement: terminal pleiochasial branches 3–5(–8), 1–4+ times 2-branched; pleiochasial bracts lanceolate or ovate to broadly ovate, wider than distal leaves; dichasial bracts distinct, broadly ovate or rhombic to triangular-ovate, base obtuse, margins entire or slightly crenate, apex obtuse, mucronate; axillary cymose branches 0–8. |
arrangement: terminal pleiochasial branches (2–)3, 5–13 cm, 3–7 times 2-branched; pleiochasial bracts ovate to oblong, 17–31 × 8–15 mm, margins entire, apex acute or obtuse; dichasial bracts ovate, 5–19 × 3–9 mm, margins entire, apex acute or obtuse; axillary cymose branches 1–5. |
Cyathia | peduncle 0.4–1 mm. |
peduncle 3–9.2 mm (often exceeding subcyathial leaves). |
2n | = 28. |
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Euphorbia brachycera |
Euphorbia telephioides |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting spring–fall. |
Habitat | Montane areas, canyons, rock crevices, sandy or gravelly slopes, pine-oak woodlands, ponderosa pine and mixed coniferous forests. | Scrubby pine flatwoods, grasslands, disturbed areas, often in sandy soils. |
Elevation | 1200–3200 m. (3900–10500 ft.) | 0–20 m. (0–100 ft.) |
Distribution |
AZ; CO; ID; MT; ND; NE; NM; SD; TX; UT; WY; Mexico (Chihuahua, Coahuila, Sonora)
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FL |
Discussion | Euphorbia brachycera is morphologically extremely variable, and plants from the northern part of the species range have sometimes been distinguished as E. robusta. Euphorbia brachycera is most closely related to the other native, perennial species of subg. Esula in western North America, namely E. chamaesula, E. lurida, E. schizoloba, and E. yaquiana. The name E. montana Engelmann, which has sometimes been applied to E. brachycera, is illegitimate because it is a later homonym of E. montana Rafinesque. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia telephioides, federally listed as threatened, is known only from Bay, Franklin, and Gulf counties in the Apalachicola region of the east-central Florida panhandle. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 300. | FNA vol. 12, p. 316. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Nummulariopsis |
Sibling taxa | ||
Synonyms | E. brachycera var. robusta, E. montana var. robusta, E. odontadenia, E. robusta, Tithymalus brachycerus, T. robustus | Galarhoeus telephioides |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) | Chapman: Fl. South. U.S., 402. (1860) |
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