Euphorbia brachycera |
Euphorbia polyphylla |
|
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horned spurge, shorthorn spurge |
lesser Florida spurge |
|
Habit | Herbs, perennial, with thick rootstock. | Herbs, perennial, with spreading rootstock. |
Stems | erect or ascending, branched, 10–60 cm, usually glabrous, sometimes puberulent. |
erect or ascending, branched, solitary, few, or occasionally densely clumped, previous year's dead stems not persistent, 18–33 cm, glabrous. |
Leaves | petiole 0–0.5 mm; blade oblong-elliptic, lanceolate, or oblanceolate to broadly ovate, 5–25 × 2–7 mm, base truncate, rounded, acute, or attenuate, margins entire, apex acute or obtuse, sometimes mucronate, surfaces usually glabrous, sometimes puberulent; venation inconspicuous, only midvein prominent. |
alternate; stipules to 0.1 mm; petiole minute or absent; blade linear to filiform, 10–20 × 0.8–1.5(–4) mm, base cuneate, margins entire, often involute, apex rounded, surfaces glabrous; venation usually obscure, midvein visible at base of wider leaves. |
Involucre | turbinate, campanulate or cupulate, 1.5–2.6 × 1.4–1.7 mm, usually glabrous, sometimes puberulent; glands 4, crescent-shaped to triangular-ovate, 0.5–0.8 × 0.7–1.7 mm, margins usually entire, occasionally slightly crenate to dentate; horns divergent, 0.2–0.4 mm, longer than teeth on gland margin. |
campanulate, 1.2–2 × 1.3–1.8 mm, glabrous or strigose on distal extreme; glands 5, red, elliptic, 0.5 × 1 mm; appendages white, orbiculate, 0.5–0.8 × 1.3–1.5 mm, erose. |
Staminate flowers | 10–15. |
20–25. |
Pistillate flower(s) | ovary, usually glabrous, sometimes puberulent; styles 0.5–0.9 mm, 2-fid. |
ovary glabrous; styles 0.6–1 mm, 2-fid at apex. |
Capsules | depressed-ovoid, 2.8–4 × 3.5–4.5 mm, 3-lobed; cocci rounded, smooth, usually glabrous, sometimes sparsely puberulent; columella 2.4–3.3 mm. |
globose, 2.3–2.8 × 4.3–5.1 mm, glabrous; columella 2.5–3.2 mm. |
Seeds | light gray, cylindric ovoid-oblong, 2–2.8(–3) × 1.4–2.2 mm, irregularly shallowly pitted; caruncle sessile to shortly stipitate, conic, 0.4–0.5 × 0.8–1.1 mm. |
ashy gray, ovoid, 2.9 × 2 mm, with obscure shallow depressions; caruncle absent. |
Cyathial | arrangement: terminal pleiochasial branches 3–5(–8), 1–4+ times 2-branched; pleiochasial bracts lanceolate or ovate to broadly ovate, wider than distal leaves; dichasial bracts distinct, broadly ovate or rhombic to triangular-ovate, base obtuse, margins entire or slightly crenate, apex obtuse, mucronate; axillary cymose branches 0–8. |
|
Cyathia | peduncle 0.4–1 mm. |
in terminal dichasia or pleiochasia; peduncle 2–6 mm (to 20 mm for central cyathium), glabrous. |
2n | = 28. |
|
Euphorbia brachycera |
Euphorbia polyphylla |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting late spring–late fall. |
Habitat | Montane areas, canyons, rock crevices, sandy or gravelly slopes, pine-oak woodlands, ponderosa pine and mixed coniferous forests. | Open sand and pine savannas. |
Elevation | 1200–3200 m. (3900–10500 ft.) | 0–10 m. (0–0 ft.) |
Distribution |
AZ; CO; ID; MT; ND; NE; NM; SD; TX; UT; WY; Mexico (Chihuahua, Coahuila, Sonora)
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FL
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Discussion | Euphorbia brachycera is morphologically extremely variable, and plants from the northern part of the species range have sometimes been distinguished as E. robusta. Euphorbia brachycera is most closely related to the other native, perennial species of subg. Esula in western North America, namely E. chamaesula, E. lurida, E. schizoloba, and E. yaquiana. The name E. montana Engelmann, which has sometimes been applied to E. brachycera, is illegitimate because it is a later homonym of E. montana Rafinesque. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia polyphylla is endemic to sandy habitats in the southern half of peninsular Florida. The species has been reported from coastal Louisiana, but whether those plants represent native occurrences or plantings is unclear (R. D. Thomas and C. M. Allen 1993–1998, vol. 2). The Florida populations are here recognized as the only native occurrences. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 300. | FNA vol. 12, p. 250. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Sibling taxa | ||
Synonyms | E. brachycera var. robusta, E. montana var. robusta, E. odontadenia, E. robusta, Tithymalus brachycerus, T. robustus | Tithymalopsis polyphylla |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) | Engelmann ex Chapman: Fl. South. U.S. ed. 2, repr. 2, 694. (1892) |
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