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horned spurge, shorthorn spurge

petty spurge

Habit Herbs, perennial, with thick rootstock. Herbs, annual, with taproot.
Stems

erect or ascending, branched, 10–60 cm, usually glabrous, sometimes puberulent.

erect, unbranched or branched, 5–30 cm, glabrous.

Leaves

petiole 0–0.5 mm;

blade oblong-elliptic, lanceolate, or oblanceolate to broadly ovate, 5–25 × 2–7 mm, base truncate, rounded, acute, or attenuate, margins entire, apex acute or obtuse, sometimes mucronate, surfaces usually glabrous, sometimes puberulent;

venation inconspicuous, only midvein prominent.

petiole 1–10 mm;

blade obovate, oblong, or suborbiculate, 5–25 × 4–15 mm, base attenuate or cuneate, margins entire, apex obtuse, surfaces glabrous;

venation pinnate, midvein prominent.

Involucre

turbinate, campanulate or cupulate, 1.5–2.6 × 1.4–1.7 mm, usually glabrous, sometimes puberulent;

glands 4, crescent-shaped to triangular-ovate, 0.5–0.8 × 0.7–1.7 mm, margins usually entire, occasionally slightly crenate to dentate;

horns divergent, 0.2–0.4 mm, longer than teeth on gland margin.

cupulate to slightly turbinate, 0.6–1.1 × 0.7–0.9 mm, glabrous;

glands 4, elliptic, 0.2–0.3 × 0.5–0.6 mm;

horns slightly convergent to divergent, 0.4–0.6 mm.

Staminate flowers

10–15.

10–15.

Pistillate flower(s)

ovary, usually glabrous, sometimes puberulent;

styles 0.5–0.9 mm, 2-fid.

ovary glabrous;

styles 0.1–0.2 mm, 2-fid.

Capsules

depressed-ovoid, 2.8–4 × 3.5–4.5 mm, 3-lobed;

cocci rounded, smooth, usually glabrous, sometimes sparsely puberulent;

columella 2.4–3.3 mm.

subglobose, 1.3–2 × 1.5–2.2 mm, slightly 3-lobed;

cocci rounded, with 2 low longitudinal wings, smooth, glabrous;

columella 1.1–1.3 mm.

Seeds

light gray, cylindric ovoid-oblong, 2–2.8(–3) × 1.4–2.2 mm, irregularly shallowly pitted;

caruncle sessile to shortly stipitate, conic, 0.4–0.5 × 0.8–1.1 mm.

whitish or grayish, subovoid, 1–1.6 × 0.6–1 mm, abaxial faces regularly large-pitted (appearing almost alveolate), adaxial faces longitudinally sulcate;

caruncle deciduous, conic, 0.2–0.5 × 0.2–0.7 mm.

Cyathial

arrangement: terminal pleiochasial branches 3–5(–8), 1–4+ times 2-branched;

pleiochasial bracts lanceolate or ovate to broadly ovate, wider than distal leaves;

dichasial bracts distinct, broadly ovate or rhombic to triangular-ovate, base obtuse, margins entire or slightly crenate, apex obtuse, mucronate;

axillary cymose branches 0–8.

arrangement: terminal pleiochasial branches 3–5, usually 2–8 times 2-branched, 1st branching level rarely 3-furcate;

pleiochasial bracts similar in shape to and slightly larger than distal leaves;

dichasial bracts distinct, ovate to obovate, base obtuse, margins entire, apex obtuse, usually mucronate;

axillary cymose branches 0–3.

Cyathia

peduncle 0.4–1 mm.

peduncle 0.4–1.1 mm.

2n

= 28.

Euphorbia brachycera

Euphorbia peplus

Phenology Flowering and fruiting spring–summer. Flowering and fruiting year-round.
Habitat Montane areas, canyons, rock crevices, sandy or gravelly slopes, pine-oak woodlands, ponderosa pine and mixed coniferous forests. Edges of gardens, weedy flower beds, roadsides, waste places, open ground near forests.
Elevation 1200–3200 m. (3900–10500 ft.) 0–1500 m. (0–4900 ft.)
Distribution
from FNA
AZ; CO; ID; MT; ND; NE; NM; SD; TX; UT; WY; Mexico (Chihuahua, Coahuila, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AZ; CA; CO; CT; DE; IA; ID; IL; IN; KY; LA; MA; MD; ME; MI; MT; NC; ND; NH; NJ; NM; NV; NY; OH; OR; PA; RI; TN; TX; UT; VA; VT; WA; WI; WV; AB; BC; MB; NB; NF; NS; ON; PE; QC; SK; Europe; w Asia [Introduced in North America; introduced also in Mexico, West Indies, Bermuda, Central America, South America, Africa, Atlantic Islands, Indian Ocean Islands, Pacific Islands, Australia]
[WildflowerSearch map]
[BONAP county map]
Discussion

Euphorbia brachycera is morphologically extremely variable, and plants from the northern part of the species range have sometimes been distinguished as E. robusta. Euphorbia brachycera is most closely related to the other native, perennial species of subg. Esula in western North America, namely E. chamaesula, E. lurida, E. schizoloba, and E. yaquiana. The name E. montana Engelmann, which has sometimes been applied to E. brachycera, is illegitimate because it is a later homonym of E. montana Rafinesque.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 300. FNA vol. 12, p. 307.
Parent taxa Euphorbiaceae > Euphorbia > subg. Esula Euphorbiaceae > Euphorbia > subg. Esula
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms E. brachycera var. robusta, E. montana var. robusta, E. odontadenia, E. robusta, Tithymalus brachycerus, T. robustus Esula peplus, Galarhoeus peplus, Tithymalus peplus
Name authority Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) Linnaeus: Sp. Pl. 1: 456. (1753)
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