Euphorbia brachycera |
Euphorbia peplus |
|
---|---|---|
horned spurge, shorthorn spurge |
petty spurge |
|
Habit | Herbs, perennial, with thick rootstock. | Herbs, annual, with taproot. |
Stems | erect or ascending, branched, 10–60 cm, usually glabrous, sometimes puberulent. |
erect, unbranched or branched, 5–30 cm, glabrous. |
Leaves | petiole 0–0.5 mm; blade oblong-elliptic, lanceolate, or oblanceolate to broadly ovate, 5–25 × 2–7 mm, base truncate, rounded, acute, or attenuate, margins entire, apex acute or obtuse, sometimes mucronate, surfaces usually glabrous, sometimes puberulent; venation inconspicuous, only midvein prominent. |
petiole 1–10 mm; blade obovate, oblong, or suborbiculate, 5–25 × 4–15 mm, base attenuate or cuneate, margins entire, apex obtuse, surfaces glabrous; venation pinnate, midvein prominent. |
Involucre | turbinate, campanulate or cupulate, 1.5–2.6 × 1.4–1.7 mm, usually glabrous, sometimes puberulent; glands 4, crescent-shaped to triangular-ovate, 0.5–0.8 × 0.7–1.7 mm, margins usually entire, occasionally slightly crenate to dentate; horns divergent, 0.2–0.4 mm, longer than teeth on gland margin. |
cupulate to slightly turbinate, 0.6–1.1 × 0.7–0.9 mm, glabrous; glands 4, elliptic, 0.2–0.3 × 0.5–0.6 mm; horns slightly convergent to divergent, 0.4–0.6 mm. |
Staminate flowers | 10–15. |
10–15. |
Pistillate flower(s) | ovary, usually glabrous, sometimes puberulent; styles 0.5–0.9 mm, 2-fid. |
ovary glabrous; styles 0.1–0.2 mm, 2-fid. |
Capsules | depressed-ovoid, 2.8–4 × 3.5–4.5 mm, 3-lobed; cocci rounded, smooth, usually glabrous, sometimes sparsely puberulent; columella 2.4–3.3 mm. |
subglobose, 1.3–2 × 1.5–2.2 mm, slightly 3-lobed; cocci rounded, with 2 low longitudinal wings, smooth, glabrous; columella 1.1–1.3 mm. |
Seeds | light gray, cylindric ovoid-oblong, 2–2.8(–3) × 1.4–2.2 mm, irregularly shallowly pitted; caruncle sessile to shortly stipitate, conic, 0.4–0.5 × 0.8–1.1 mm. |
whitish or grayish, subovoid, 1–1.6 × 0.6–1 mm, abaxial faces regularly large-pitted (appearing almost alveolate), adaxial faces longitudinally sulcate; caruncle deciduous, conic, 0.2–0.5 × 0.2–0.7 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3–5(–8), 1–4+ times 2-branched; pleiochasial bracts lanceolate or ovate to broadly ovate, wider than distal leaves; dichasial bracts distinct, broadly ovate or rhombic to triangular-ovate, base obtuse, margins entire or slightly crenate, apex obtuse, mucronate; axillary cymose branches 0–8. |
arrangement: terminal pleiochasial branches 3–5, usually 2–8 times 2-branched, 1st branching level rarely 3-furcate; pleiochasial bracts similar in shape to and slightly larger than distal leaves; dichasial bracts distinct, ovate to obovate, base obtuse, margins entire, apex obtuse, usually mucronate; axillary cymose branches 0–3. |
Cyathia | peduncle 0.4–1 mm. |
peduncle 0.4–1.1 mm. |
2n | = 28. |
|
Euphorbia brachycera |
Euphorbia peplus |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting year-round. |
Habitat | Montane areas, canyons, rock crevices, sandy or gravelly slopes, pine-oak woodlands, ponderosa pine and mixed coniferous forests. | Edges of gardens, weedy flower beds, roadsides, waste places, open ground near forests. |
Elevation | 1200–3200 m. (3900–10500 ft.) | 0–1500 m. (0–4900 ft.) |
Distribution |
AZ; CO; ID; MT; ND; NE; NM; SD; TX; UT; WY; Mexico (Chihuahua, Coahuila, Sonora)
|
AL; AZ; CA; CO; CT; DE; IA; ID; IL; IN; KY; LA; MA; MD; ME; MI; MT; NC; ND; NH; NJ; NM; NV; NY; OH; OR; PA; RI; TN; TX; UT; VA; VT; WA; WI; WV; AB; BC; MB; NB; NF; NS; ON; PE; QC; SK; Europe; w Asia [Introduced in North America; introduced also in Mexico, West Indies, Bermuda, Central America, South America, Africa, Atlantic Islands, Indian Ocean Islands, Pacific Islands, Australia]
|
Discussion | Euphorbia brachycera is morphologically extremely variable, and plants from the northern part of the species range have sometimes been distinguished as E. robusta. Euphorbia brachycera is most closely related to the other native, perennial species of subg. Esula in western North America, namely E. chamaesula, E. lurida, E. schizoloba, and E. yaquiana. The name E. montana Engelmann, which has sometimes been applied to E. brachycera, is illegitimate because it is a later homonym of E. montana Rafinesque. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
|
Source | FNA vol. 12, p. 300. | FNA vol. 12, p. 307. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | E. brachycera var. robusta, E. montana var. robusta, E. odontadenia, E. robusta, Tithymalus brachycerus, T. robustus | Esula peplus, Galarhoeus peplus, Tithymalus peplus |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) | Linnaeus: Sp. Pl. 1: 456. (1753) |
Web links |
|