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horned spurge, shorthorn spurge
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| Herbs, perennial, with thick rootstock. |
Herbs, annual, with taproot. |
erect or ascending, branched, 10–60 cm, usually glabrous, sometimes puberulent. |
erect-ascending, often basally decumbent, often branched near base, 12–28 cm, glabrous. |
petiole 0–0.5 mm; blade oblong-elliptic, lanceolate, or oblanceolate to broadly ovate, 5–25 × 2–7 mm, base truncate, rounded, acute, or attenuate, margins entire, apex acute or obtuse, sometimes mucronate, surfaces usually glabrous, sometimes puberulent; venation inconspicuous, only midvein prominent. |
petiole 0–3 mm; blade broadly oblanceolate to subspatulate, or proximalmost often orbiculate, 3–20 × 3–9 mm, base attenuate, margins entire, apex rounded to emarginate, surfaces glabrous; venation pinnate, midvein prominent. |
turbinate, campanulate or cupulate, 1.5–2.6 × 1.4–1.7 mm, usually glabrous, sometimes puberulent; glands 4, crescent-shaped to triangular-ovate, 0.5–0.8 × 0.7–1.7 mm, margins usually entire, occasionally slightly crenate to dentate; horns divergent, 0.2–0.4 mm, longer than teeth on gland margin. |
infundibular, 1.3–1.6 × 1–1.2 mm, glabrous; glands 4, crescent-shaped; 0.4–0.5 × 0.8–1 mm; horns divergent, 0.2–0.4 mm. |
10–15. |
15–20. |
ovary, usually glabrous, sometimes puberulent; styles 0.5–0.9 mm, 2-fid. |
ovary glabrous; styles 0.8 mm, 2-fid. |
depressed-ovoid, 2.8–4 × 3.5–4.5 mm, 3-lobed; cocci rounded, smooth, usually glabrous, sometimes sparsely puberulent; columella 2.4–3.3 mm. |
ovoid-globose, 2.6–2.7 × 2.5–2.7 mm, slightly lobed; cocci rounded to ± flattened, smooth, glabrous; columella 2–2.1 mm. |
light gray, cylindric ovoid-oblong, 2–2.8(–3) × 1.4–2.2 mm, irregularly shallowly pitted; caruncle sessile to shortly stipitate, conic, 0.4–0.5 × 0.8–1.1 mm. |
dark brown, oblong-ovoid, 1.5–2 × 1–1.6 mm, with deep, rounded pits in 3–4 regular vertical rows; caruncle reniform-ovate, conic, 0.5–0.7 × 0.5–0.8 mm. |
arrangement: terminal pleiochasial branches 3–5(–8), 1–4+ times 2-branched; pleiochasial bracts lanceolate or ovate to broadly ovate, wider than distal leaves; dichasial bracts distinct, broadly ovate or rhombic to triangular-ovate, base obtuse, margins entire or slightly crenate, apex obtuse, mucronate; axillary cymose branches 0–8. |
arrangement: terminal pleiochasial branches 3, 1–3 times 2-branched; pleiochasial bracts ovate-deltate to subrhombic-ovate, shorter and wider than distal leaves; dichasial bracts connate 1/4 length, not imbricate, broadly deltate to subreniform, base truncate to broadly obtuse, margins entire, apex obtuse to bluntly acuminate; axillary cymose branches 1–5. |
peduncle 0.4–1 mm. |
peduncle 0.4–0.7 mm. |
= 28. |
= 26. |
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| Flowering and fruiting spring–summer. |
Flowering and fruiting spring. |
| Montane areas, canyons, rock crevices, sandy or gravelly slopes, pine-oak woodlands, ponderosa pine and mixed coniferous forests. |
Semiopen forests, bluffs and ledges, stream banks, glades. |
| 1200–3200 m. (3900–10500 ft.) |
100–400 m. (300–1300 ft.) |
|
AZ; CO; ID; MT; ND; NE; NM; SD; TX; UT; WY; Mexico (Chihuahua, Coahuila, Sonora)
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AR; MO; OK; TN; TX |
Euphorbia brachycera is morphologically extremely variable, and plants from the northern part of the species range have sometimes been distinguished as E. robusta. Euphorbia brachycera is most closely related to the other native, perennial species of subg. Esula in western North America, namely E. chamaesula, E. lurida, E. schizoloba, and E. yaquiana. The name E. montana Engelmann, which has sometimes been applied to E. brachycera, is illegitimate because it is a later homonym of E. montana Rafinesque. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia ouachitana is restricted to semiopen forests and woodlands in the south-central United States. It is similar to E. commutata and has been most commonly identified as that species in the past. However, it differs in its consistently brown seeds that have pits in regular, vertical rows. The two species also differ in E. commutata’s numerous, proximal, long-petiolate elliptic leaves. The proximal leaves of E. ouachitana are spatulate with an orbiculate blade and petiolelike base. Aside from the restricted distribution of E. ouachitana in Missouri and Tennessee, the ranges of E. ouachitana and E. commutata do not overlap (see M. H. Mayfield 2013 for a detailed discussion of the distribution). Euphorbia ouachitana is most common in the Ouachita Mountains from southeastern Oklahoma to Hot Springs County, Arkansas. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
| FNA vol. 12, p. 300. |
FNA vol. 12, p. 306. |
| Euphorbiaceae > Euphorbia > subg. Esula |
Euphorbiaceae > Euphorbia > subg. Esula |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
| E. brachycera var. robusta, E. montana var. robusta, E. odontadenia, E. robusta, Tithymalus brachycerus, T. robustus |
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| Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) |
Mayfield: J. Bot. Res. Inst. Texas 7: 642, figs. 2[row 2, right], 6. (2013) |
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