Euphorbia brachycera |
Euphorbia exigua |
|
---|---|---|
horned spurge, shorthorn spurge |
dwarf spurge, small spurge |
|
Habit | Herbs, perennial, with thick rootstock. | Herbs, annual, with taproot. |
Stems | erect or ascending, branched, 10–60 cm, usually glabrous, sometimes puberulent. |
erect, unbranched or branched, 3–30 cm, glabrous. |
Leaves | petiole 0–0.5 mm; blade oblong-elliptic, lanceolate, or oblanceolate to broadly ovate, 5–25 × 2–7 mm, base truncate, rounded, acute, or attenuate, margins entire, apex acute or obtuse, sometimes mucronate, surfaces usually glabrous, sometimes puberulent; venation inconspicuous, only midvein prominent. |
petiole 0–0.5 mm; blade linear, linear-oblong, or linear-spatulate, 2–30 × 1–5 mm, base cuneate or truncate, margins entire, apex acute, obtuse, or emarginate, surfaces glabrous; venation inconspicuous, only midvein prominent. |
Involucre | turbinate, campanulate or cupulate, 1.5–2.6 × 1.4–1.7 mm, usually glabrous, sometimes puberulent; glands 4, crescent-shaped to triangular-ovate, 0.5–0.8 × 0.7–1.7 mm, margins usually entire, occasionally slightly crenate to dentate; horns divergent, 0.2–0.4 mm, longer than teeth on gland margin. |
cupulate, 0.3–0.5 × 0.6–0.8 mm, glabrous; glands 4, elliptic to crescent-shaped, 0.2–0.4 × 0.3–0.6 mm; horns divergent, 0.2–0.5 mm. |
Staminate flowers | 10–15. |
5–8. |
Pistillate flower(s) | ovary, usually glabrous, sometimes puberulent; styles 0.5–0.9 mm, 2-fid. |
ovary glabrous; styles 0.5–0.7 mm, 2-fid. |
Capsules | depressed-ovoid, 2.8–4 × 3.5–4.5 mm, 3-lobed; cocci rounded, smooth, usually glabrous, sometimes sparsely puberulent; columella 2.4–3.3 mm. |
subglobose, 1–1.8 × 1.3–2 mm, slightly lobed; cocci rounded, smooth, puncticulate toward abaxial line, glabrous; columella 1.1–1.4 mm. |
Seeds | light gray, cylindric ovoid-oblong, 2–2.8(–3) × 1.4–2.2 mm, irregularly shallowly pitted; caruncle sessile to shortly stipitate, conic, 0.4–0.5 × 0.8–1.1 mm. |
blackish to grayish, 4-angled-ovoid, 1–1.5 × 0.5–0.7 mm, white, tuberculate; caruncle conic or subconic, 0.1–0.3 × 0.1–0.3 mm. |
Cyathial | arrangement: terminal pleiochasial branches 3–5(–8), 1–4+ times 2-branched; pleiochasial bracts lanceolate or ovate to broadly ovate, wider than distal leaves; dichasial bracts distinct, broadly ovate or rhombic to triangular-ovate, base obtuse, margins entire or slightly crenate, apex obtuse, mucronate; axillary cymose branches 0–8. |
arrangement: terminal pleiochasial branches 3–5, each 1–3 times 2-branched; pleiochasial bracts similar in shape and size to or sometimes slightly longer and wider than distal leaves; dichasial bracts distinct, linear, or linear-lanceolate, or lanceolate-ovate, base rounded to subcordate, margins entire, apex acute; axillary cymose branches 0–5. |
Cyathia | peduncle 0.4–1 mm. |
peduncle 0–2 mm. |
2n | = 28. |
|
Euphorbia brachycera |
Euphorbia exigua |
|
Phenology | Flowering and fruiting spring–summer. | Flowering and fruiting summer–fall. |
Habitat | Montane areas, canyons, rock crevices, sandy or gravelly slopes, pine-oak woodlands, ponderosa pine and mixed coniferous forests. | Edges of gardens, roadsides, waste places. |
Elevation | 1200–3200 m. (3900–10500 ft.) | 0–1500 m. (0–4900 ft.) |
Distribution |
AZ; CO; ID; MT; ND; NE; NM; SD; TX; UT; WY; Mexico (Chihuahua, Coahuila, Sonora)
|
CA; NY; WV; BC; NS; ON; Europe; w Asia; n Africa; Atlantic Islands (Macaronesia) [Introduced in North America] |
Discussion | Euphorbia brachycera is morphologically extremely variable, and plants from the northern part of the species range have sometimes been distinguished as E. robusta. Euphorbia brachycera is most closely related to the other native, perennial species of subg. Esula in western North America, namely E. chamaesula, E. lurida, E. schizoloba, and E. yaquiana. The name E. montana Engelmann, which has sometimes been applied to E. brachycera, is illegitimate because it is a later homonym of E. montana Rafinesque. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Euphorbia exigua can be easily distinguished from other annual species of the genus in the flora area by its tuberculate seeds. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
Source | FNA vol. 12, p. 300. | FNA vol. 12, p. 302. |
Parent taxa | Euphorbiaceae > Euphorbia > subg. Esula | Euphorbiaceae > Euphorbia > subg. Esula |
Sibling taxa | ||
Synonyms | E. brachycera var. robusta, E. montana var. robusta, E. odontadenia, E. robusta, Tithymalus brachycerus, T. robustus | Tithymalus exiguus |
Name authority | Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 192. (1859) | Linnaeus: Sp. Pl. 1: 456. (1753) |
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