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black-seed spurge
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desert spurge, Sonoran sandmat, tiny sandmat
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| Herbs, annual, with slender taproot. |
Herbs, annual, with slender taproot. |
erect, branched, 10–35 cm, glabrous or strigillose (especially when young and around nodes). |
prostrate, mat-forming, 5–35 cm, glabrous or shortly pilose. |
opposite proximally, alternate distally; stipules 0.1–0.2 mm; petiole 1–4(–6) mm, glabrous, sericeous or strigillose; blade linear to narrowly elliptic, 8–52 × 2–7 mm, base attenuate, margins entire, ciliate-strigose, apex acute, abaxial surface sparsely strigillose to sericeous, adaxial surface usually glabrous; venation obscure, only midvein conspicuous. |
opposite; stipules distinct, subulate, 0.2–0.4 mm, pilose; petiole 0.5–1.2 mm, glabrous or pilose; blade ovate to elliptic, 6–15 × 2–4 mm, base asymmetric, one side cuneate to rounded, other side rounded, margins entire, apex obtuse, surfaces glabrous or pilose; venation obscure or only midvein conspicuous. |
obconic, 0.9–1.5 × 0.9–1.3 mm, strigillose to pilose; glands 5, yellow or pink, U-shaped, 0.2–0.3 × 0.4–0.5 mm; appendages greenish, white, or pink, forming narrow rim around gland, or ovate, oblong, or obovate and usually 2-fid, rarely rudimentary, 0.2–1.4 × 0.2–0.6 mm, entire. |
campanulate, 0.4–0.6 × 0.5–0.9 mm, glabrous or pilose; glands 4, red, circular to oblong, 0.1 × 0.1–0.2 mm; appendages absent. |
20–25. |
2–5. |
ovary glabrous, puberulent, strigillose, or pilose; styles 0.5–0.8 mm, 2-fid 1/3–1/2 length. |
ovary usually glabrous, rarely pilose; styles 0.1–0.2 mm, 2-fid at apex. |
oblate, 1.5–2.6 × 2.1–3.3 mm, glabrous or puberulent, strigillose, or pilose; columella 1.2–2.1 mm. |
oblong, 1.3–1.5 × 1.1–1.3 mm, usually glabrous, rarely pilose; columella 1–1.2 mm. |
brown to grayish black, narrowly ovoid, 3- or 4-angled in cross section, sometimes obscurely so, 1.3–1.9 × 1–1.4 mm, tuberculate, often with shallow depressions; caruncle absent. |
light gray, narrowly ovoid, 4-angled in cross section, 0.9–1 × 0.5–0.6 mm, smooth to slightly rugose or with 1–4 faint transverse ridges that do not pass through abaxial keel. |
solitary at distal nodes or in weakly defined cymes or dichasia, dichasial bracts and distal stem leaves wholly green; peduncle 0.5–3.6 mm, strigillose. |
solitary at distal nodes; peduncle 0.4–1.4 mm. |
= 32. |
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| Flowering and fruiting spring–fall. |
Flowering nearly year-round in response to sufficient rainfall. |
| Sandy and rocky soils on slopes and canyon bottoms in pine-juniper woodlands, oak woodlands, grasslands. |
Desert scrub, riparian woods with ash and willow, saltbush scrub, Joshua tree woodlands and grasslands, often in sandy or gravelly areas. |
| 1400–2600 m. (4600–8500 ft.) |
-20–1800 m. (-100–5900 ft.) |
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AZ; NM; TX; Mexico (Chihuahua, Durango, Sonora)
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AZ; CA; NM; NV; TX; UT; Mexico (Baja California, Baja California Sur, Chihuahua, Coahuila, Durango, Sonora)
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In Texas, Euphorbia bilobata is known only from Jeff Davis County. (Discussion copyrighted by Flora of North America; reprinted with permission.) |
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| FNA vol. 12, p. 243. |
FNA vol. 12, p. 278. |
| Euphorbiaceae > Euphorbia > sect. Alectoroctonum |
Euphorbiaceae > Euphorbia > sect. Anisophyllum |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana |
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Chamaesyce micromera |
| Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 190. (1859) |
Boissier: Proc. Amer. Acad. Arts 5: 171. (1861) |
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