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black-seed spurge

graceful sandmat, graceful spurge

Habit Herbs, annual, with slender taproot. Herbs, annual, with taproot.
Stems

erect, branched, 10–35 cm, glabrous or strigillose (especially when young and around nodes).

erect to ascending, 15–50 cm, glabrous.

Leaves

opposite proximally, alternate distally;

stipules 0.1–0.2 mm;

petiole 1–4(–6) mm, glabrous, sericeous or strigillose;

blade linear to narrowly elliptic, 8–52 × 2–7 mm, base attenuate, margins entire, ciliate-strigose, apex acute, abaxial surface sparsely strigillose to sericeous, adaxial surface usually glabrous;

venation obscure, only midvein conspicuous.

opposite;

stipules connate, deltate, usually entire, sometimes laciniate-fringed at tip, 1.5–2.2 mm, glabrous;

petiole 1–3 mm, glabrous;

blade obliquely oblong-oblanceolate, 10–35 × 7–15 mm, base asymmetric, oblique, margins serrate or serrulate, especially toward apex, apex broadly acute, surfaces glabrous; palmately veined at base, pinnate distally.

Involucre

obconic, 0.9–1.5 × 0.9–1.3 mm, strigillose to pilose;

glands 5, yellow or pink, U-shaped, 0.2–0.3 × 0.4–0.5 mm;

appendages greenish, white, or pink, forming narrow rim around gland, or ovate, oblong, or obovate and usually 2-fid, rarely rudimentary, 0.2–1.4 × 0.2–0.6 mm, entire.

obconic, 0.9–1.1 × 0.4–0.9 mm, glabrous;

glands 4, yellow-green to brown, stipitate, subcircular, 0.2 × 0.2 mm, occasionally nearly rudimentary;

appendages absent on smaller glands or white to pink, shape highly variable, usually round to ± elliptic, 0.3–0.4 × 0.5–0.7 mm, distal margin entire.

Staminate flowers

20–25.

(0–)2–20.

Pistillate flowers

ovary glabrous, puberulent, strigillose, or pilose;

styles 0.5–0.8 mm, 2-fid 1/3–1/2 length.

ovary glabrous;

styles 0.4 mm, 2-fid 1/2 length.

Capsules

oblate, 1.5–2.6 × 2.1–3.3 mm, glabrous or puberulent, strigillose, or pilose;

columella 1.2–2.1 mm.

depressed-globoid, 1.3–1.4 × 1.1–1.5 mm, glabrous;

columella 1–1.1 mm.

Seeds

brown to grayish black, narrowly ovoid, 3- or 4-angled in cross section, sometimes obscurely so, 1.3–1.9 × 1–1.4 mm, tuberculate, often with shallow depressions;

caruncle absent.

with very thin whitish mucilaginous coat over light brown testa below, ovoid-triangular, bluntly 4-angled in cross section, 0.9–1.1 × 0.5 mm, with shallow irregular depressions alternating with low, smooth ridges.

Cyathia

solitary at distal nodes or in weakly defined cymes or dichasia, dichasial bracts and distal stem leaves wholly green;

peduncle 0.5–3.6 mm, strigillose.

in dense, axillary and terminal, capitate glomerules with reduced, bractlike leaves subtending cyathia;

peduncle 0.5–1.8 mm.

2n

= 32.

Euphorbia bilobata

Euphorbia hypericifolia

Phenology Flowering and fruiting spring–fall. Flowering and fruiting early spring–late fall.
Habitat Sandy and rocky soils on slopes and canyon bottoms in pine-juniper woodlands, oak woodlands, grasslands. Open, disturbed areas, nurseries.
Elevation 1400–2600 m. (4600–8500 ft.) 0–200 m. (0–700 ft.)
Distribution
from FNA
AZ; NM; TX; Mexico (Chihuahua, Durango, Sonora)
[WildflowerSearch map]
[BONAP county map]
from FNA
AL; AR; FL; GA; LA; OK; SC; TX; Central America; South America; West Indies [Introduced in Asia, Pacific Islands]
[WildflowerSearch map]
[BONAP county map]
Discussion

In Texas, Euphorbia bilobata is known only from Jeff Davis County.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Euphorbia hypericifolia is native to the New World tropics, and it is most likely adventive in the flora area (where it is most widely distributed in Florida and Texas). Reports from Arizona, California, and Maryland likely represent waifs or misidentifications.

(Discussion copyrighted by Flora of North America; reprinted with permission.)

Source FNA vol. 12, p. 243. FNA vol. 12, p. 272.
Parent taxa Euphorbiaceae > Euphorbia > sect. Alectoroctonum Euphorbiaceae > Euphorbia > sect. Anisophyllum
Sibling taxa
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hypericifolia, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
E. aaron-rossii, E. abramsiana, E. acuta, E. agraria, E. albomarginata, E. alta, E. angusta, E. antisyphilitica, E. arizonica, E. astyla, E. austrotexana, E. bicolor, E. bifurcata, E. bilobata, E. blodgettii, E. bombensis, E. brachycera, E. capitellata, E. carunculata, E. chaetocalyx, E. chamaesula, E. cinerascens, E. commutata, E. conferta, E. cordifolia, E. corollata, E. crenulata, E. cumulicola, E. cuphosperma, E. curtisii, E. cyathophora, E. cyparissias, E. davidii, E. deltoidea, E. dendroides, E. dentata, E. discoidalis, E. eriantha, E. exigua, E. exserta, E. exstipulata, E. falcata, E. fendleri, E. florida, E. floridana, E. garberi, E. georgiana, E. geyeri, E. glyptosperma, E. golondrina, E. gracillima, E. graminea, E. helioscopia, E. helleri, E. heterophylla, E. hexagona, E. hirta, E. hooveri, E. humistrata, E. hyssopifolia, E. indivisa, E. innocua, E. inundata, E. ipecacuanhae, E. jaegeri, E. jejuna, E. laredana, E. lasiocarpa, E. lata, E. lathyris, E. longicruris, E. lurida, E. macropus, E. maculata, E. marginata, E. meganaesos, E. melanadenia, E. mendezii, E. mercurialina, E. mesembrianthemifolia, E. micromera, E. misera, E. missurica, E. myrsinites, E. nephradenia, E. nutans, E. oblongata, E. ocellata, E. ophthalmica, E. ouachitana, E. parishii, E. parryi, E. pediculifera, E. peplidion, E. peplus, E. perennans, E. pergamena, E. pinetorum, E. platyphyllos, E. platysperma, E. polycarpa, E. polygonifolia, E. polyphylla, E. porteriana, E. prostrata, E. pubentissima, E. purpurea, E. radians, E. rayturneri, E. revoluta, E. roemeriana, E. rosescens, E. schizoloba, E. serpens, E. serpillifolia, E. serrata, E. serrula, E. setiloba, E. simulans, E. spathulata, E. stictospora, E. strictior, E. telephioides, E. terracina, E. tetrapora, E. texana, E. theriaca, E. thymifolia, E. tithymaloides, E. trachysperma, E. trichotoma, E. vallis-mortae, E. velleriflora, E. vermiculata, E. villifera, E. virgata, E. wrightii, E. yaquiana
Synonyms Chamaesyce glomerifera, C. hypericifolia, E. glomerifera
Name authority Engelmann: in W. H. Emory, Rep. U.S. Mex. Bound. 2(1): 190. (1859) Linnaeus: Sp. Pl. 1: 454. (1753)
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